640 UMBELLIFERAE 



Open stony hills, 'JO to G800 i'eet : ISicn-a Nevada from Kern Co. to Lassen Co.; 

 Coast Ranges from Kern Co. to Siskiyou Co. North to Washington, east to 

 Colorado. Apr.-May, fr. May-July. 



Locs. — Sierra Nevada: Greenhorn Eange, Kern Co., Hall 4" Bahcock 5076; Alder Creek, 

 Yosemite, Jepson 4318 ; Mariposa, Congdon ; Cold Spr., Tuolumne Co., Jepson 6460 ; Honey Lake, 

 T. Brandegee ; Olinda, Shasta Co., Blankinship. South Coast Ranges: San Emigdio Canon, sw. 

 Kern Co., Davy 2071; Jolon, Monterey Co., Eastwood; Pico Blanco, Santa Lucia Mts., Davy 

 7332; Coyote sta., Santa Clara Valley, Jepson 14,297; Redwood City, Jepson 5736, 5738; Mt. 

 Diablo, Jepson 3120; Antioch, Curran. North Coast Ranges: Sonoma Valley, Jepson 4184; 

 Vandcn, Solano Co., Jepsnn 14,809 ; Wcldon Canon, Vaca Mts., Jepson 7197 ; Howell Mt. foot- 

 hills. Jepson 14,384; Asa Bean Ridge, ne. Mendocino Co., Jepson 14,295; South Yollo BoUy, se. 

 Trinity Co., Jepson 14,296; Crane Creek, w. Tehama Co., Jepson 14,294; Devils Backbone, sw. 

 Siskiyou Co., Jepson 2103 ; Yreka, Butler 744. 



Var. douglasli Jepson. Leaf -segments larger, as much as 4^/^ lines long and 1 line wide; 

 umbel sometimes with a single bract. — California, Douglas. 



Var. ellipticum Jepson. Root-crowTi bearing short leafy branches; leaf -segments oblong, 

 cuspidate; fruit broadly elliptical, notched at base and apex, 9^4 lines long and 4^^ lines broad, 

 the wings twice as wide as the body; dorsal and intermediate ribs obscure or filiform. — Feather 

 River near Marysville, Bigelow; region of the "head waters of the Sacramento" (Pit River 

 branch), Snyder. 



Refs. — LoMATiUM MACROCARPUM C. & R. Contrib. U. S. Nat. Herb. 7:217 (1900) ; Jepson, 

 Man. 721 (1925). Peucedanum macrocarpum Nutt. ; T. & G. Fl. 1:627 (1840), type loc. Colum- 

 bia River, Ore., Nuttall; Jepson, Fl. W. Mid. Cal. 358 (1901), ed. 2, 302 (1911). Cogswellia 

 macrocarpa Jones, Contrib. W. Bot. 12:33 (1908). Var. douglasii Jepson, Madroiio 1:153 

 (1924), type from Cal., Douglas; Man. 722 (1925). Var. ellipticum Jepson, Madrono 1:153 

 (1924), Man. 722 (1925). Peucedanum nudicaule var. ellipticum T. & G. Pac. R. Rep. 2:121 

 (1855), type loc. "Round Valley near the sources of the Sacramento", that is, on Pit River branch, 

 Snyder. Lomatium ellipticum C. & R. I.e. 7:217 (1900). 



9. L. dasycarpum C. & R. Lace Parsnip. Plants 1 to l^^ feet high, the 

 peduncles arising from the root-crown or from very short stems; herbage puberu- 

 lent or pubescent, peduncles and wings of carpels often purplish; leaves ternately 

 decompound and dissected into oblong or linear segments, these 1 to 2 or 3 lines 

 long; umbels somewhat equally 6 to 15-rayed; rays 1 to 2 inches long; bractlets 

 linear to ovate, more or less tomentose; pedicels in fruit about I/2 inch long, usually 

 longer than the carpels; flowers white; fruit purplish or whitish, broadly elliptic 

 to orbicular, with subcordate base, 3i/^ to 4 (or 7) lines long, the wings thin-mem- 

 branaceous, 2 to 3 times the width of the very narrow or somewhat spindle-shaped 

 body; body commonly very woolly when young, more or less glabrate in age; oil- 

 tubes usually 1 in the intervals (often a second one in the lateral intervals), 2 to 

 4 on the commissure. 



Dry hillsides and mesas, 200 to 2000 (or 4000) feet: coastal Southern Califor- 

 nia; South Coast Ranges from San Benito Co. to Contra Costa Co. Feb. -Apr., fr. 

 May-June. 



Note on variation. — The prevailing plant of the South Coast Ranges is here taken as typical 

 of the Douglas type of Lomatium dasycarpum. This usual form has the ultimate leaf-segments 

 oblong to broadly linear and % to 1 line long, and the oil-tubes commonly 1 in the intervals. The 

 reddish wings of the fruit develop early, are only slightly pubescent and thus differ markedly from 

 the tomentulose or pubescent body. The number of oil-tubes is not, however, constant, usually not 

 even in a single individual, and hence is not a critical differentia. It will thus be seen from the 

 specimens cited below that, as we proceed northward in the South Coast Ranges, the oil-tubes are 

 more frequently 2 or 3 in the intervals, 1 oil-tube being less commonly found. The plants of the 

 North Coast Ranges have most commonly 2 or 3 oil-tubes in the intervals ; in addition the bractlets 

 are usually more strongly developed (mostly broadly ovate) and the body of the carpel is broader, 

 nearly equaling the wings in breadth. These northern plants we cite under var. medium Jepson. 

 In habit, foliage, pedicels, in size of fruit and thinness of wings the var. medium is more closely 

 associated with L. dasycarpum than -with L. tomentosum, although in character of the oil-tubes 

 it may be regarded as intermediate between these two species. As here accepted L. tomentosum 

 is restricted geographically to the area of the Great Valley and Sierra Nevada foothills. It has 

 linear or filiform leaf -segments (lYz to 2 lines long) and larger fruits (6 to 7 lines long) ; the 

 ovary is densely woolly, but by the time it is half grovwi the wings are often still inconspicuous, so 

 tardily do they develop. 



