PARSLEY FAMILY 649 



1. A. hendersonii C. & R. Coast Angelica. Very stout, I/2 to 3 feet high; 

 herbage more or less tomentose, especially on the inflorescence and under surface 

 of the leaves ; leaves quinate or ternate, then pinnate ; leaflets tending to be crowded, 

 thick, broadly ovate, 1^/2 to 2^ inches long, obtuse (or rarely acute), serrate ; um- 

 bels rather condensed; rays in flower subequal; fruiting rays % to 2^4 inches long; 

 umbellets dense, as if capitate; pedicels 1 line long or less; bractlets 1 or 2, linear- 

 acuminate; fruit broadly oblong, slightly pubescent, 3 to 4 lines long; lateral wings 

 thick and corky, as broad as the body; oil-tubes solitary in the intervals; seed deeply 

 sulcate beneath the oil-tubes. 



Sea-bluffs or flats along the coast, 5 to 300 feet : San Francisco Co. to Del Norte 

 Co. North to Washington. July-Aug. 



Phylogenetic note. — The leaves are a little fleshy and rigid ; the pinnae stand at right angles 

 to the rachis, the leaflets at right angles to the secondary rachis. The plants are somewhat squat 

 or dwarfish and the leaves are condensed as is the inflorescence. In all these particulars Angelica 

 hendersonii seems as if it were, phylogenetically, a littoral adaptation of the inland A. tomentosa 

 Wats. 



Locs.— Pt. Lobos, San Francisco (Fl. W. Mid. Cal. 356); Pt. Eeyes, Davy 6869; Salmon 

 Creek, Sonoma coast, Jepson 15,940 ; Newport, Mendocino Co., Jepson 13,487 ; Loleta, Humboldt 

 Co., Jepson 2133 ; Crescent City, E. Paris 3393. 



Eefs. — Angelica hendersonii C. & E. Bot. Gaz. 18 :80 (1888), type loc. Long Beach, Ilwaco, 

 Wash., Eenderson 2158; Jepson, Fl. W. Mid. Cal. 355 (1901), ed. 2, 299 (1911), Man. 727, fig. 

 711 (1925). 



2. A. tomentosa Wats. Wood Angelica. Stout, 2 to 8 feet high, the stems 

 and especially the leaves pubenilent, or sometimes nearly glabrous; leaves bipin- 

 nate or ternate or quinate and then pinnate ; leaflets ovate, acute, acutish or often 

 long-pointed, sometimes varying to lanceolate or roundish, irregularly serrate, ob- 

 liquely 2-lobed, or not lobed and merely oblique, l^i to 3 (or 6 ) inches long; petioles 

 strongly dilated at base; fruiting rays 1 to 5 inches long; fruiting pedicels 2 to 3 

 lines long; ovary tomentulose; fruit oblong or elliptic, glabrous, 3 to 4% lines long; 

 dorsal and intermediate ribs small and acutish ; lateral wings nearly equaling the 

 body in breadth; oil-tubes 1 in the intervals, or sometimes 2 in the lateral intervals, 

 mostly 2 on the commissure ; seed somewhat sulcate beneath the oil-tubes. 



Shady woods, 200 to 7000 feet : coastal Southern California; Coast Ranges from 

 Santa Cruz Co. to Humboldt Co. June-July. 



Locs. — Coastal S. Cal.: Palomar Mt., McClatchie ; Mt. San Jacinto (Univ. Cal. Publ. Bot. 

 1:98); Waterman Canon, San Bernardino Mts., Parish; West Fork Cucamonga Canon, San 

 Gabriel Mts. ; Cow Canon, San Gabriel Eiver, Peirson 2508 ; Big Pine Mt., Santa Barbara Co., 

 J. B. Hall. Coast Eanges: Santa Cruz Mts. (Anderson, Nat. Hist. Santa Cruz Co. 38) ; Crystal 

 Springs Lake, San Mateo Co., C. F. BaJcer 3354 ; Berkeley, H. A. Walker; Mt. Tamalpais, Jepson; 

 St. Helena, Jepson 13,483 ; Calistoga, Jepson 13,484; Peanut, Trinity Co., J. W. Patton; Eureka, 

 Tracy 6902 ; Alton, Humboldt Co., Tracy 6560. 



Var. californica Jepson. Bays very unequal (1^ to 5% inches long), scaberulous at the 

 ends; oil-tubes 2 (or 3) in the intervals, mostly 4 in lateral pairs on the commissure. — Vaca Mts. 



Eefs. — Angelica tomentosa Wats. Proc. Am. Acad. 11 : 141 (1876), type loc. San Francisco ; 

 Jepson, Fl. W. Mid. Cal. 356 (1901), ed. 2, 300 (1911), Man. 728 (1925). A. tomentosa var. 

 elata Jepson, Fl. W. Mid. Cal. 356 (1901), ed. 2, 300 (1911), type loc. Napa Valley, Jepson. Var. 

 CALIFORNICA Jepson, Fl. W. Mid. Cal. 356 (1901), ed. 2, 300 (1911), Man. 728 (1925). A. cali- 

 fornica Jepson, Erythea 1:8 (1893), type loc. Gates Canon, Vaca Mts., Jepson 14,246, 14,250. 



3. A. breweri Gray. Sierra Angelica. (Fig. 270.) Stems 3 to 5 feet high; 

 herbage glabrous; leaves ternate or quinate, then pinnate; leaflets lanceolate to 

 oblong or occasionally ovate-lanceolate, acuminate, sharply serrate, 1% to 4 inches 

 long; rays many, II/2 to 2i/^ inches long in fruit; pedicels 2 to 3^ lines long; rays 

 and pedicels not "web-footed" or very obscurely so; pedicels and ovaries whitish- 

 puberulent; fruit oblong or somewhat narrowed below, pubescent or becoming 

 glabrous, 3 to 41/2 lines long ; dorsal and intermediate ribs more or less prominent ; 

 lateral wings as wide as the body; oil-tubes 1 or 2 in the intervals. 



