556 PAPAVERACEAE 



8p. (/) Sometimes we have very interesting specimens which bear on the same plant both 

 monilifomi carpels with rounded beads and carpels in which the beads are short-cylindric 

 instead of rounded, all seed-bearing: San Diego (ocean cliffs), K. Brandegee Ip. Cf. P. 

 splMerocarpus Greene, (g) In other cases the fruiting cai'pels are unusually long (9 to 11 

 lines), the carpels each with an obvious dorsal keel: Sutter plains, Jepson; Antioch to Marsh 

 Creek, near the coal mines, K. Brandegee a. Cf. P. emarginatiis Greene, (h) The fruiting 

 carpels are sometimes t-\risted and either sterile or fertile: Ocean View, San Francisco, K. 

 Brandegee 12p, 13p, 14p, 15p. Cf. P. ccmtorius Greene, based on a spm. from Lake Co., or 

 Colusa Co., Curran. (i) There is frequently a tendency to extreme hairiness of carpels. This 

 variation seems correlated with extreme hairiness of stems and buds, especially when young: 

 Springville, Tulare Co., Purpns 1749. On the other hand the reverse correlation is not true, 

 since Purpus 5000 (Erskin Greek, Kern Co.) has ver}' hairy buds and stems, but glabrous 

 carpels. 



It is possible that further collections and field studies may show some of the specific segre- 

 gates, listed below as reductions, to be of varietal rank, but it is more likely that accumulation 

 of knowledge concerning the range and kind of variation will destroy this possibility. No other 

 Californian species seems so inviting for a study of fluctuating variability, particularly in 

 intensive garden cultures. It is probable that such work would reveal the most unexpected 

 surprises. 



Locs. — Coast Ranges: Dinsmore Banch, near Buck Mt, Humboldt Co.. Tracy 4139; Fort 

 Seward Ridge, Humboldt Co., Jepson 1902 (3000 ft.); Cahto, Mendocino Co., T. Brandegee; 

 Snow Mt., Lake Co., T. Brandegee; Sites. Colusa Co., K. Brandegee: St. Helena, Jepson 6240; 

 Yountville, Jepson; Berkeley, Jepson; Moraga Valley, Jepson; Crystal Springs Lake, San 

 Mateo Co., C. F. Baker 433. Great Valley: Crane Creek, Salt Creek and Wimmeshaw, w. 

 Tehama Co., Jepson; Chieo, HcUer 10718; Sutter plains, Jepson; Wilson Creek, Vacaville, 

 Jepson; Tracy, C. F. Baker 2780; Madera, Davy 1697. Sierra Nevada: Weber Creek, Eldorado 

 Co., K. Brandegee ; Phoenix Lake, Tuolumne Co., A. L. Grant .50; Greenliorn Range, Hall 4' 

 Babcock 5016. Southern California: Swarthout Caiion, San Antonio Mts., Hall 1520; Colton, 

 Parish; Thomas Valley, San Jacinto Mts., Hall 543; Ramona, Jepson 8514; San Felipe Valley, 

 e. San Diego Co., Jepson 8734. 



Var. crinitus Greene. Peduncles and buds with longer hairs, often densely pilose; buds 

 globose; petals yellow, often with pink or greenish tips — Tehachapi Mts. south to the Cuya- 

 maca Mts.: Teh'achapi, Greene; Coahuilla VaUey, Riverside Co., Jepson 1470. This variety 

 is not definitely different from the species but merely represents a terminus of a line of 

 variation. 



Var. horrldulus Jepson n. comb. Petals spreading nearly rotately from a turbinate or 

 almost cylindric base 3 to 4 lines long; carpels white when young vnth a dense covering of 

 stifRsh hairs, the hairs rapidly deciduous and the carpels soon becoming only thinly hairy. — 

 Southern Sierra Nevada foothills from Fresno Co. to Tulare Co., 3500 to 5000 ft.: ridge n. 

 of Pinehurst, Fresno Co., Ottley 1438, Newlon 202; e. of Springville, Tulare Co., Purptts 1749. 

 The following spms. have cai-pels destitute of the heavy coat of hairs just described, being in 

 pubescence similar to the ordinary form of Platystemon californicus, but they have turbinate 

 corollas in various degrees or intergrades: Pleasant Caiion, Panamint Mts., Hall 4' Chandler 

 6958; Erskine Creek, Kern Co., Pjirp^ts 5000; Leonis Valley, betw. Manz.ana and Gorman. 

 Davy 2659. These facts evidence that the corolla character is more constant than the 

 pubescence character, although the latter is the more striking. In those plants (as first cited 

 above) where both characters are found intensified we have a state which is really an extreme 

 in a series of variations. 



Var. nutans Brandegee. Fmits nodding. — Coast of Southern California; San Diego, T. 

 Brandegee : Santa Cruz Isl., T. Brandegee. 



Refs. — Platystemon californicus Benth. Trans. Hort. Soc. Lond. ser. 2, 1:405 (1835), 

 type from Cal., Douglas; Jepson, Fl. W. Mid. Cal. 205 (1901). Var. crinitus Greene, Fl. Fr. 

 282 (1892). P. crinitns Greene, Pitt., 2:13 (1889), type loc. Tehachapi, Greene. Var. hor- 

 RIDULUS Jepson. P. liorridtdiis Greene, Pitt. 5:178 (1903), type loc. betw. Sequoia and Sanger 

 Mills, Fresno Co., Eastwood. Var. nutans Br.andg. Zoe 5:177 (1903), type loo. San Diego, 

 T. Brandegee. P. nw/on,? Greene, Pitt. 5 :192 (1903). 



The following additional segregates are here alphabetically arranged: P. aculeolatus Greene, 

 Pitt. 5:167 (1903), type loc. Santa Barbara Isl., Trask; depressed, about 5 inches broad; 

 corolla \i inch broad (ex. char.). P. acutatvs Greene I.e. 187, type loc. Middle Tule River, 

 Purptos; Fedde in Engler, Pfizr. 4"':12fi, fig. 18A-C (1909). P. dnemonoides Greene, I.e. 177, 

 type loc. Alcalde, w. Fresno Co., Eastwood. P. antoninus Greene I.e. 180, type loc. San 

 Antonio River, Santa Lucia Mts., Eastwood. P. arvorum Greene I.e. 174, tvpe loc. Tracy, C. F. 

 Baker 3199. P. califortiictis var. sphaeroearpus T. Brandegee. Zoe 5:177 (1903), type loc. 

 Colusa Jet., T. Brandegee. P. capsnlaris Greene I.e. 165, type loc. San Simeon, T. Brandegee; 

 Fedde I.e. 116, fig. 15c. P. ccrinvninis Greene I.e. 169, type loc. San Rafael, J. P. Moore; Fedde 

 I.e. 117, fig. 15e a-c; var. styloii(.s Greene I.e. 170, based on spms. from San Francisco Co., 

 Kellogg 4' Harford, and Redwood Caiion, Marin Co., Michener 4- Bioletti; Fedde 1.0. 117, fig. 



