374 



TREMANDRACE^. 



[HypoCtYnous Exogens. 



Order CXXXII. TREMANDRACE^.— Poreworts. 



Tremandi-acese, R. Brown in Flinders, p. 12, (1814) ; DC. Prodr. 1. 343. (1824) ; Endl Gen. ccxxxii. 



Diagnosis. — Sapindal Exoyens, with partially eom2ilete symmetrical jloicers, a valvate 

 calyx^ and 2-A-ceUed anthers opening hy pores. 

 Slender heath-like slu'ubs, with theu' han-s usually glandular. Leaves alternate or 

 whorled, %\-ithout stipules, entu'e or toothed. Pedicels 

 solitary, axillary, 1 -flowered. Flowers often large and 

 showy. Sepals 4 or 5, equal, with a valvate aestivation, 

 slightly cohermg at the base, and deciduous. Petals 

 equal in number to the sepals, ^\ith an involute aestiva- 

 tion, wrapping up the stamens in pau's, much larger than 

 the calyx, and deciduous. Stamens hypogjTious, distinct, 

 2 before each petal, and therefore either 8 or 10 ; anthers 

 2- or 4-celled, opening by a pore at the apex. Ovaiy 

 2-celled ; o\Tiles from 1 to 3 in each cell, anatropal, with 

 a hooked apex, pendulous ; styles 1 or 2. Fruit capsular, 

 2-celled, 2-valved ; dehiscence locuhcidal. Seeds pendu- 

 lous, ovate, with a hooked appendage at the apex, but 

 with none about the liiliun ; embryo cylindrical, straight, 

 in the axis of fleshy albumen, and about half as long, the 

 radicle next the liilum. 



There is little to diN^de these plants from Milkworts, 

 except their regular sjonmetrical flowers, and valvate 

 calyx. They want the caruncula of that Order, in room 

 of which they have the chalazal end of the seed extended 

 into a hooked process. Theii' stamens being opposite the 

 petals in pau's may, taken \Ai\\ the valvate calyx, be 

 regarded as an indication of some tendency towards 



Rhamnads, and the general condition of theu' flower is much Uke that of Pittosporads, 



except in the great development of the embryo. 

 All are natives of New Holland. 

 Theu* properties are vmknown. 

 De CandoUe placed them between Milkworts and Pittosporads, Meisner between 



Frankeniads and IVIilkworts, Endlicher in his Polygalinae consisting of Poreworts and 



Milkworts only, and Adolphe Brongniart takes the same view of theu* affuiity. 



Fig CCLX. 



GENERA. 



Tetratheca, Sm. 

 Tremandra, R. Br. 

 Platytheca, Steetz. 



Numbers. Gex. 3. Sp. 16. 



TiliacecB. 



Position. Tremandrace^. — Polygalacese. 



Pittosporacca;, 



Fig. CCLX.— Tetratheca hirsuta. 1. the stamens ; 2. the pistD, with one of the cells laid open. 



