names published before 1921, but not treated by Safford 
at that time, see Sachet (10) and Fosberg (7). Collections 
of several species are very limited or absent in a number 
of our larger herbaria (A, COL, GH, NY, US). 
Of the numerous collections of conspicuous and highly 
attractive tree Daturas in these herbaria, nearly all are 
referable to three taxa: D. candida (Pers.) Saff., D. 
suaveolens H. & B. ex Willd. and D. sanguinea R. & P. 
DeWolf (6) and the herbarium determinations of A. S. 
Barclay in 1959 are in agreement here. Each of the re- 
maining collections most frequently displays a unique 
combination of characters, and several such collections 
have been selected for the typification of new species. 
In 1955, Schultes (12) described as Methysticodendron 
A mesianum what I believe to be a greatly modified tree 
Datura from the Valley of Sibundoy in southern Colom- 
bia. 
It is interesting to note that, in 1891, Wettstein’s 
summary (15) of the Solanaceae for Engler and Prantl’s 
Die Natiirlichen Pflanzenfamilien cited only three arbo- 
rescent Daturas, although several more had been de- 
scribed by that time. He cited D. sanguinea, D. suaveo- 
lens and D. arborea, the last very probably referring to 
plants identified here as D. candida, and thus he antici- 
pated by more than half a century the growing realization 
(3, 5, 6) that the principal species of tree Daturas are 
few in number. 
The very extensive work of Blakeslee and his associates 
(1) with the herbaceous Daturas demonstrated a great 
range of variability and the spontaneous appearance of 
many unusual characteristics. Of the 541 gene mutations 
encountered, 72 appeared following heating, wounding 
and ageing, or spontaneously in nature. Recessive genes 
controlling leaf shape, flower size, shape and color, and 
fruit form are among those uncovered. It is entirely 
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