can be regarded as an incipient species still confined to 
a single locality in southern Colombia. 
Cotomsia: Barclay & Schultes 147 (Tver), 149, 177; Pennell 7093; 
Pérez-Arbeldez & Cuatrecasas 5960; von Sneidern 1898. 
D. dolichocarpa (Lagerh.) Safford in Journ. Wash. 
Acad. Sci. 11: 186 (1921), based upon two collections 
by Sodiro from Ecuador, is distinguished chiefly by its 
very long (81 cm.), terete fruit. It resembles D. swaveo- 
lens in the very long corolla, stamens and pistil, but the 
short calyx, long corolla teeth and free anthers relate it 
to D. candida. Bristol 1387 (GH), from one of Sodiro’s 
two collecting localities (Santo Domingo de los Colora- 
dos), has a fruit intermediate in size (16.5 em.) and shape 
between this species and D. candida, but an extremely 
long corolla (44.3 em.) and calyx (28 cm.) much exceed- 
ing those of D. dolichocarpa which are, respectively, 35 
em. and 13cm. The calyces of this collection have con- 
spicuous horn-like apices, as in J. cornigera Hooker. 
D. longifolia (agerh.) Safford ibid. 186 (1921) is 
closely allied to D. dolichocarpa, but the single Ecua- 
dorean collection by Sodiro is described as having very 
long, linear-oblong and sinuate-repand leaves, thus ap- 
proaching those of Methysticodendron Amesianum R. K. 
Schultes, a striking clone discussed below. The style is 
also unusually long. 
D. versicolor (Lagerh.) Safford ibid. 183 (1921) also 
closely resembles D. dolichocarpa but has a pointed calyx, 
ultimately reddish corolla, shorter stamens and pistil, a 
very long aristate fruit, and thicker, verrucose seeds. 
Four of the tree Datura concepts are best interpreted 
as hybrids, three of them between D. candida and D. 
sanguinea. 
D. arborea Linnaeus Sp. Pl. 1: 179 (1758), the first 
to be described, is rarely seen. While its affinity is with 
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