phase bridges as noted above in Table III. ‘The remain- 
der might well be caused by the presence of genetic fac- 
tors for pollen abortion; Blakeslee and his co-workers 
found thirty such factors in their extensive cultures of 
herbaceous Daturas (1). 
Flower variation among the Datura candida cultivars 
All of the cultivars have markedly similar flowers, with 
the exception of the bizarre ‘Culebra’ which is discussed 
later. Nevertheless, each displays a distinctive range of 
variation in the length of the several floral parts. Plate 
L. compares nine of the cultivars according to the lengths 
of five of their floral parts. It is evident that with but 
few exceptions the ranges of variation of each cultivar 
are either wholly within, or largely overlapping, those 
of D. candida ‘Buyés’. This strongly suggests that most 
of the cultivars were selected from the large and variable 
population of D. candida ‘Buyés’. It further suggests 
that the Sibundoy did not select and propagate new 
forms (cultivars) characterized by floral differences, but, 
as we shall see below, they selected leaf differences for 
preferential maintenance as clones. 
Fruit variation 
Most of the fruits found in prolonged searching 
throughout the Valley were on ‘Buyés’ and ‘Amaron’ 
trees which are by far the most numerous. Some fruits 
were also seen and collected on ‘Dientes’, ‘Quinde’ and 
‘Biangiin’ trees; the remainder of the D. candida culti- 
vars are believed by the natives never to form fruit. The 
indehiscent fruits eventually dry up, and the pericarp 
disintegrates slowly: but the seeds are not released, and 
the dried fruit often remains on the tree for several 
months. Plate LI shows the variation in fruit size and 
shape among the five cultivars known (however rarely) 
[176 ] 
