Finally, we must mention the important monograph of Robyns 
(1931); after comparative studies on floral organization in the 
Solanaceae, he verified and extended the early observations of 
Wydler and Eichler concerning the pattern of symmetry of the 
several verticils. Of his various conclusions, it is worthwhile to 
quote the following (1931: 79): “L’organisation florale des Sola- 
nacées ne peut donc se comprendre qu-en relation avec lobliquité 
du gynécée, quien est le caractere fondamental et distinctif et qui 
a produit le type bilabié propre a la famille. Le Solanacées for- 
ment une famille homogéne et naturelle, associant un type vege- 
tatif particulier a un type floral tout aussi caractéristique et dont 
toute l’évolution s est faite d apres un plan uniforme, sous Vaction 
de la tendance a la zygomorphie, qui s‘est manifestée dans toutes 
les tribes”. 
GENERIC DELIMITATION AND AFFINITIES 
It is surprising that Bentham (1845: 143) proposed Leptoglossis 
without any mention of Sa/piglossis asa closely related genus. He 
compared it instead with Schwenckia L.and Browallia L., genera 
which are quite different. Nevertheless, the important point Is 
that Sa/piglossis, Leptiglossis and Revesia were recognized as 
separate generic entities by Bentham. On the contrary, Wettstein 
(1891: 36) included Leproglossis and Revesia as sections in Salpi- 
glossis.,Since then taxonomists in general have accepted Wett- 
stein’s classification (Werdermann, 1928; Johnston, 1928, 1929; 
Millan, 1931), until Hunziker (1977: 45) reinstated the three 
genera on the basis of morphological and phytogeographical 
evidence. D’Arcy (1978) also favored this opinion. 
The affinities of each genus will be explained further on in this 
paper. Since all there genera have been confused at one time or 
the other with Browallia L. and Nierembergia R. et P. (Torrey, 
1859: 155, 156; Hemsley 1882: 437), or with Hunzikera D’Arcy 
(Gray, 1877: 164; Millan, 1941: 489) or Bouchetia Dun. in DC. 
(D’Arcy, 1978: 714), the following artificial key is presented to 
facilitate their identification: 
’The delimitation between Sa/piglossideae and Nicotianeae is not clear; further studies are 
needed for a proper understanding of both tribes. 
