Erythroxylum Coca var. Ipadu Plowman var. nov. 
Ab Erythroxylo Coca typico ramis fortiter erectis virgatis, 
ramentis paucis vel nullis, foliis apice rotundatis, lineis abaxiali- 
bus obscuris, pedicellis brevioribus, stamineo urceolo valde 10- 
denticulato, stigma promienti, ovato-oblonga differt. 
TYPE: PERU: Dept. Loreto: Prov. Maynas. Rio Ampiyacu, Puca Urquillo and 
vicinity, approx. lat. 3°05’, long. 71°55’. Cultivated shrub 2.5 m tall in open 
coca plantation in jungle clearing, interplanted with yuca and fruit trees. 
Branches slender, erect, then arching, leaves mostly near the tips, bright green. 
Flowers cream. N.v. jibina (Witoto). 5 Apr 1977. T. Plowman, R. E. Schultes 
& O. Tovar 6663 (holotype, ECON; isotypes, F 1824462, GH, K, MO, S, U, 
US, USM). 
Slender shrub to 3 m. tall. Single trunk to 2 cm. in diameter. 
Bark light greyish brown with fine longitudinal cracks. Branches 
strongly erect, spindly, sometimes arching over with age. Branch- 
lets straight, not knobby at the nodes, light yellowish green 
becoming reddish brown, with lenticular or elongated lenticels. 
Ramenta inconspicuous, sparse or sometimes absent, if present 
found only at the base of new shoots. Leaves present mostly at 
the branch tips, the blade elliptic to broadly elliptic or elliptic- 
ovate, apically obtuse to rounded, basally obtuse to acute, some- 
times briefly attenuate, 35-115 mm long, 20-42 mm wide, firmly 
membranaceous, dark to medium green above, pale glaucous 
green beneath, dull to somewhat shiny above, dull beneath, mid- 
rib yellowish green, with an acute adaxial ridge, lateral nerves 
often prominulous above, abaxial longitudinal lines obscure or 
wanting, central panel often faint or concolorous. Petiole 2—4 
mm. long. Flowers in axils of past season’s growth, usually on 
naked stems but sometimes axillary in subterminal leaf axils, 
1-5(6) per node, scattered or sometimes crowded, creamy white 
to yellowish throughout, odorless. Pedicel short, thickened dis- 
tally, 2-4 mm long, to 1.5 mm in diameter. Lamina of petals 
spreading, sometimes suberect and not fully opening, incurved 
at apex. Margin of staminal cup markedly 10-denticulate, the 
‘The term “ramenta” was first used by Martius (1843) for the persistent, often congested. 
scale-like structures which frequently cover the twigs of Erythroxylum species. Mor- 
phologically, ramenta are simply stipules which are produced without accompanying 
leaves. 
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