teeth short, triangular. Short-styled flowers predominant, style 
short 1.5-2 mm. long, stigma large, ovate-oblong, | mm long, 
0.5 mm broad, bright yellowish green. Drupe 7-8 mm long, 4-6 
mm in diameter, embryo and endosperm usually aborted. 
Chromosome number n = 12. 
ETYMOLOGY: This variety is named for the Brazilian vernacular 
name of Amazonian coca. 
COMMON NAMES: idapu, yvpadu (Amazonian Brazil), coca 
(Amazonian Peru), jibina (Witoto, Peru), (pi (Bora, Peru), patu 
(Cubeo, Colombia), botd (Maku, Brazil). 
DISTRIBUTION: Cultivated in the Amazon basin of Peru, Co- 
lumbia and Brazil, along the Amazon River and its major 
tributaries. 
SPECIMENS STUDIED: 
BRAZIL: AMAZONAS: Basin of Rio Ica, near San Antonio, “padu” or “coca”, 
Dec 1935, Krukoff 7645 (NY); Rio Purus basin, Rio Uneiuxi, Maku Indian 
village, 300 km above mouth, “bot6” (Maku), 23 Oct 1971, Prance et al. 15572 
(ECON, INPA, K, NY, U, US); Rio Jurua, Marary, Sep 1900, Ule 5039 (K, L); 
Rio Solimées, Tefe, Lago de Tejé, Mujeira, “padu”, 13 Jul 1973, Lieras et al. 
P16681 (GH); Tefe, “padu”, “ipadu”, 16 Jul 1972, PLK & Urbana 12231 
(INPA); Tefe, Fazenda Experimental, 20 Feb 1973, PLK & Marilene 12553 
(INPA); “prope Ega et Sao Paulo d’Olivencga”, 1819-1820, Martius s.n. (M); 
“in sylvis Japurensibus”, “ypadu”, 1819, Martius S.N. (M); Ilha Parauary, 20 
Oct 1874, Traill 77 (K); Rio Negro, Providentia, “ipadu”, Jul 1888, Prinzessin 
Therese von Bayern s.n. (M); Rio Negro, Cucui, 7.5.1973, Silva et al. 1310 (F); 
Rio Janauari, “paida”, Spruce 73 (K): Manaus, igarape do Buiao, “ipadu”, 
5-7-1958, Coélho s.n. (F). PARA: Belem, “ipadu”, Sep-Oct 1961, Pires 51919 
(NY, US). [Belem, Sao Jozé to the Arsenal, 1828-1830], Burchell 9588 (K), 
Belém, Botanical Garden, Museu Goeldi, 15 May 1908, Baker 70 (A, BO, GH, 
LE, MO, NY, US); Rio Guama, Apr 1899, E. Poisson s.n. (P); Santarem, May 
1929, Dahlgren & Sella 36 (F). 
COLOMBIA: PUTUMAYO: Rio Putumayo, entre Puerto Asis y Puerto Legu- 
isamo, 300-400 m, 14-15 Oct 1954, Garcia-Barriga et al. 18713 (COL). 
AMAZONAS: Leticia, camino a Tarapaca, km. 17, 18 Jul 1965, Lozano & 
Ospina 488 (COL); Tikuna village, km. 12 on trail from Leticia to La Pedrera, 
14 Apr 1975, Cabrera 3367(COL). Rio Igaraparana, La Chorrera, 180 m, 6 Jun 
1942, Schultes 3899(ECON); Ro Igaraparana, “kudu jibina” (Witoto), 1 Mar 
1974, Idrobo 6859 (COL, ECON); Rio Igaraparana, 15 km. abajo de La Chor- 
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