lateral nerves and veinlets usually obscure above, rarely promin- 
ulous, abaxial longitudinal lines and central panel usually 
inconspicuous or obscure. Petiole 1-5 mm. long. Flowers in 
axils of previous season’s growth, often near the tips of the 
branchlets or sometimes among the persisting leaves, usually 
1-3(10) per node, scattered, if congested then only briefly, witha 
strong, foetid odor resembling raw pumpkin. Pedicels long, 3-11 
mm. long, median length 9 mm. Petals fugaceous or sometimes 
persisting, appearing rotate, 3.5-5 mm. long, 2-3 mm. wide, the 
lamina subcymbiform or the midrib depressed above, yellowish 
green to cream, ligule creamy white. Staminal cup half the 
length to equaling the calyx. Drupe ovoid to ellipsoid, some- 
times fusiform, obtuse to acute at apex, 10-13 mm. long, 4-7 
mm. in diameter. Whole plant, especially the leaves, suffused 
with odor of wintergreen (methyl salicylate). Chromosome 
number 2n = 24. 
ETYMOLOGY: The variety is named for the city of Trujillo in 
northern Peru, near the area of primary cultivation and from 
which the leaves are exported. 
COMMON NAMES: coca de Trujillo, tupa (Peru): coca (Ecuador). 
DISTRIBUTION: Cultivated in northern Peru on the western 
slopes of the Andes and in the valley of the Alto Rio Maranon. 
Disjunct populations known from northwestern Ecuador and 
adjacent Colombia. 
SPECIMENS STUDIED: 
COLOMBIA: NARINO: Rio San Juan, across river from Maldonado, 2000 m., 
“coca”, 26 Jan 1977, Boeke 854 (F); Tumaco, 9 May 1926, O.F. Cook 98 (US). 
ECUADOR: CARCHI: Maldonado, 1450-1650 m., “coca”, 2 June 1978, Madi- 
son etal. 4920 (F, SEL); Environs of Chical, 12 km. below Maldonado, along 
Rio San Juan, 1200 m., “coca”, 25 May 1978, Madison et al. 4447 (F, NCU, 
SEL). 
PERU: AMAZONAS: Prov. Chachapoyas: Rio Utcubamba, km. 6 from La 
Caclid, 1600 m., 25 Feb 1976, Plowman 5563 (ECON, F, USM), Plowman 
5564 (ECON, F, K, USM), Plowman 5565 (ECON, F, NCU, RB, USM). 
Yumbay, above Balsas near Hornopampa, alt. 1100 m, 27 Feb 1976, Plowman 
5583 (ECON, F, K, NCU, US, USM), Plowman 5587 (ECON, F, USM), 
Plowman 5588 (ECON, F, NCU, USM), Plowman 5589 (ECON, F, USM), 
Plowman 5590 (ECON, F, K, USM). 
Prov. Contumaza: Dist. Simbron, Fundo Farrat, 8 Jan 1966, Machado 2960 
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