TABLE 1 (continued) 
Flower 
Calyx circumscissile near the 
base and falls away with the 
corolla, the persistent base ulti- 
mately forming a membranous 
disk, cup or reflexed frill sub- 
tending the mature fruit. 
Fruit a relatively small, dehis- 
cent berry or capsule borne on 
short pedicels in an erect, sub- 
erect or nodding position. 
Bicarpellate and tetralocular due 
to presence of false septa. 
Fruit in most species possessing 
a dehiscence mechanism. 
Pericarp usually spinose. 
Seeds relatively small and lack- 
ing a corky seed coat. 
Seeds usually with a well de- 
Calyx not circumscissile, either 
falling away entirely or forming 
a persistent husk-like structure 
around the mature fruit. 
Fruit a large pendulous berry 
borne on much elongated pedi- 
cels. 
Bicarpellate and bilocular owing 
to the lack of false septa. 
Fruit lacking any dehiscence 
mechanism. 
Pericarp smooth and unarmed. 
Seeds large and most species 
with a thick, corky seed coat. 
Seeds lacking a caruncle. 
veloped funicular caruncle. 
to hummingbird pollination, and fruits that are large and 
fleshy. The derived nature of these characters would 
seem to preclude the possibility of Brugmansia being 
ancestral to Datura. 
The unique combination of characters of Datura cera- 
tocaula has caused Bernhardi (8), Safford (17) and others 
to consider it a connecting link between Brugmansia and 
Datura. The plant is an herbaceous annual with erect 
flowers, a circumscissile calyx, a tetralocular ovary, and 
‘arunculate seeds. hese are all characters typical of 
Datura. It differs from other species of Datura, how- 
ever, in having a smooth, fleshy fruit and from all other 
| O77 | 
