numerous in the larger intercostal areas of E. novogranateme 

 var. novogranatense. Intramarginal veins of secondary origin 

 are absent from leaves of all varieties. 



Slender tertiary veins arise at widely acute to slightly obtuse 

 angles (80-100°) along the admedial (inner) and exmedial 

 (outer) flanks of the secondary veins and ramify within the 

 mtercostal areas to form non-oriented, polygonal, tertiary 

 areoles of various sizes. Tertiary areoles tend to be slightly 

 smaller and most numerous in both varieties of E. Coca 

 Conversely, leaves of E novogranateme var. novogranatense 

 typically have fewer but larger intercostal areas and tertiary 

 areoles than do those of either variety oi E Coca. Leaves of £. 

 novogranatense var. truxiUense are often intermediate in this 

 respect but may reveal the full range of patterns exhibited 

 collectively among leaves of the other three varieties (Plate 40). 

 All tertiary areoles are subdivided by a random reticulum of 

 slender, fourth and fifth order veinlets, which constitute a 



irately well developed system of small, non-oriented, 

 polygonal and quadrangular areoles. This system of areoles is 

 best developed (i.e. most completely closed) and contains more 

 numerous areoles in both varieties of £. Coca, whereas leaves of 



mod 



norma 



of fewer and slightly larger areoles (Plates 39 and 40). 



The relative number of freely terminating veinlets in coca 

 leaves serves as a quantification of qualitative differences in high 

 order venation and areolation among the four cultivated 

 varieties. Leaves of E. Coca varieties often contain fewer veinlet 



m 



than do those of either variety of E. novogranatense (Table 5- 

 Plates 39 and 40). Although leaves of E novogranatense var' 

 truxiUense have the highest veinlet termination numbers of all 

 coca varieties, they may show a degree of variation equal to that 

 observed collectively among the other three varieties, as is the 

 case with qualitative aspects of their form and venation (Table 

 5). Veinlet termini are typically tracheoid with spiral or pitted 

 secondary walls (Plate 40). Tracheoid idioblasts were not 

 observed in any variety. 



The marginal ultimate venation of all coca leaves consists of 



309 



