their leaf structure, the archeological leaves studied here are 

 easily distinguished from any wild "coca mimics" (Rury, 1981, 

 1982). 



Features shared by archeological and recent coca leaves 

 include their: (1) size and form; (2) variable prominence of a 

 central panel on the lower leaf surface; (3) patterns of low and 

 high order venation; (4) comparable numbers of freely terminat- 

 ing veinlets per mm^ of leaf area; (5) polygonal epidermal cells 

 with straight (anticlinal) walls; (6) epidermal papillae restricted 

 to the lower leaf surface; (7) paracytic stomata of similar 

 dimensions and numbers, also restricted to the lower leaf 

 surface; and (8) numerous prismatic crystals associated with 

 veins and occurring within subdivided epidermal cells. 



All archeological coca leaves studied here are morphologically 

 identical to recent leaves of Trujillo coca {E. novogranatense 

 var. truxillense) in their small size, mostly lanceolate shape, 

 acute leaf apices and cuneate leaf bases, as well as their reduced 

 prominence of a central panel. As in recent Trujillo coca, the 

 archeological leaves reveal a basic foliar venation pattern which 

 is intermediate between the characteristic eucamptodromous 

 configuration of E. Coca var. Coca and the slender-veined, 

 brochidodromous pattern of most leaves of E. novogranatense 

 var. novogranatense. Details of leaf venation in the archeo- 

 logical leaves which conform to our typology for Trujillo coca 

 include: (1) the divergence angles and irregular course of the 

 secondary veins; (2) an intermediate number of secondary veins 

 and intercostal areas; (3) a relatively incomplete (open) system 

 of high order venation, with (4) a rather large number of freely 



terminating veinlets. 



Statistical details of Erythroxylum leaf structure are both 

 ecologically more variable and taxonomically less reliable than 

 qualitative aspects of foliar form, venation and anatomy. 

 Nevertheless, our archeological leaves fall clearly within the 

 range of statistical variation in stomatal and venation systems 

 observed among leaves of all four modern coca varieties (Table 

 5). The archeological leaves are more similar in these features to 

 both varieties of E. novogranatense than to either variety of E. 

 Coca, but most closely resemble leaves of Trujillo coca in their 



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