THE NEWTS STOMACH DURING DIGESTION 425 



Celis lining Ihe surface of Ihe slomach, 



In stud3'ing the changes that occur during and after sécrétion in the 

 cells lining the surface of the stomach great difficulties are encountered, 

 because thèse cells exhibit in an}- one spécimen so many différent phases, 

 that it becomes almost impossible in some cases to say exactly the condi- 

 tion which corresponds to a given hour; it has therefore been found neces- 

 sary to describe as the typical condition at each hour the appearances pre- 

 sented by the greatest number of cells. 



I propose first to describe briefly the appearances of such a cell in the 

 resting condition and then to point out the changes as they occur hour by 

 hour as in the case of the oxyntic cells. 



Resting cells. 



The cells are long and narrow somewhat pyramidal or conical in shape 

 with rounded free extremities. They average 40 — ■ 50 v- in length and are 

 arranged side by side in a single row. The free end of the cell is occupied 

 by a clear, elongated, mucigen-holding cup with rounded extremities and 

 straight sides. This cup measures some 12 y. in length and 9 i^ in breadth. 

 In spécimens stained with methyl blue eosine it appears of a pale sky blue 

 colour owing to the présence in it of very many rounded granules of mucigen 

 that exhibit a feeble affinity for the methyl blue. When stained by my me- 

 thod already detailed, the granules become bright reddish violet and are 

 then readily seen lying in the meshes of a wide meshed spongioplasm, 

 which is most apparent around the margins and lowest part of the cup. 

 The free surface of the cup is covered by a perforated cell wall, the minute 

 pores in which are well spaced and can be readily seen in many cells. 



Immediately beneath the cup, between it and the nucleus and stret- 

 ching outwards to the junction of the cup with the cell wall, there is a 

 zone of protoplasm in which minute knotted thread-like structures running 

 vertically can be distinguished. Thèse cords extend inwards to the level of 

 the middle of the nucleus, where they appear to terminate in the netlike 

 spongioplasm. The hyaloplasm between there is filled with very minute 

 granules that stain vividly with eosine as described by Langley'^ This zone 

 varies somewhat in thickness ; when measured from the upper extremity of 

 the nucleus to the most inward extcnding part of the cup averages 3,5 — 



