no. S630 CLEPTORIA — HOFFMAN 27 



and robust, with a massive and convex solenomerite, and the sternal 

 processes of the third segment are the most prominent in the genus. 

 Obvious affinity is seen with C. rileyi, which has a somewhat more 

 slender telopodite, and with C. bipraesidens, in which the solenomerite 

 is longer and not notably thickened. These three species might be 

 considered as making up a single group. Cleptoria macro, is intermediate 

 between the members of this group and C. divergens. On the diagram 

 (fig. 22), the presumed sequence of specialization runs from right to 

 left. 



A second diagram (fig. 23) shows the remarkably good "fit" that 

 obtains when the foregoing derivation is superimposed on the present 

 distribution of the various species. Although species of Sigmoria now 

 occur widely in the South Carolina Piedmont and may be sympatric 

 with C. macro, at the present, the majority of species of that genus 

 occur in the Southern Appalachians. It seems reasonable to suspect 

 that the ancestral lineage of Cleptoria developed in that area and 

 subsequently migrated south and west through the Piedmont of 

 Georgia. There is some indication that speciation may have occurred 

 in a sort of straight-line (perhaps "leap-frog") sequence rather than 

 by in situ speciation from a single widespread ancestral form. 



Literature Cited 



Bollman, Charles H. 



1S88. Descriptions of fourteen new species of North American myriapods. 

 Proc. U.S. Nat. Mus., vol. 10, pp. 617-627. 

 Chamberlin, Ralph V. 



1939. On some diplopods of the family Fontariidae. Bull. Univ. Utah, 

 Biol. Ser., vol. 5, no. 3, pp. 1-19, figs. 1-37. 

 Chamberlin, Ralph V., and Hoffman, Richard L. 



1958. Checklist of the millipeds of North America. U.S. Nat. Mus. Bull. 

 212, 236 pp. 

 Hoffman, Richard L. 



1950. Records and descriptions of diplopods from the southern Appalach- 

 ians. Journ. Elisha Mitchell Sci. Soc, vol. 66, pp. 11-33, figs. 1-32. 

 Loomis, H. F. 



1943. New cave and epigaean millipeds of the United States, with notes on 



some established species. Bull. Mus. Comp. Zool., vol. 92, no. 7, 

 pp. 371-410, figs. 1-18. 



1944. Millipeds principally collected by Professor V. E. Shelford in the 



eastern and southeastern United States. Psyche, vol. 51, pp. 

 166-177, figs. 1-6. 



US. GOVERNMENT PRINTING OFFICE: 1967 



