no. 3635 MONOPHYLLUS — SCHWARTZ AND JONES 7 



moved. The major difference between M. clinedaphus and the other 

 "species" recognized in the same paper — M. redmani, M. portori- 

 censis, M. plethodon — was that the "plane of the basioccipital and 

 basisphenoid, instead of sloping forward at a faint angle as in the other 

 species of the genus, pitches abruptly forward and downward at an 

 angle of 32°." It may be recalled also that Miller had no material at 

 that time from Cuba or Hispaniola and, thus, was unaware of the size 

 and general characters of populations from those islands. 



Gary L. Ranck (pers. comm., Feb. 8, 1966) has taken measurements 

 for us of the skull and skin of the holotype (USNM 5210/37405) of M. 

 clinedaphus. Pertinent measurements are: forearm 39.6; greatest 

 length of skull 21.7, condylobasal length 19.6, postorbital constriction 

 4.1, maxillary toothrow 7.8. Ranck also confirmed the peculiarity of 

 the basioccipital-presphenoid angle on which Miller placed so much 

 emphasis, but Ranck suggested (as had C. O. Handley previously) that 

 it may well be due to the fact that the skull was removed from a speci- 

 men in spirits and that preparation and subsequent drying possibly 

 resulted in distortion of the skull. Later, the junior author examined 

 the skull of M. clinedaphus and reached a similar conclusion, i.e., that 

 it somehow had been distorted after collection. The base of the brain- 

 case, just above the basioccipital-presphenoid angle, clearly indicates 

 that the skull has been bent, probably while wet and, therefore, pos- 

 sibly prior to the time it was extracted for study. Additionally, the 

 palate is damaged, having been laterally compressed so that the tooth- 

 rows are closer together than usual and nearly parallel. In any event, 

 in all measurements except condylobasal length the holotype of M. 

 clinedaphus agrees with Monophyllus examined from Cuba (the con- 

 dylobasal length in the holotype is 19.6, whereas the lowest measure- 

 ment in nine Cuban bats is 19.7). In the maxillary toothrow, the first 

 two premolars are separated by a diastema slightly more than half the 

 length of the first tooth, indicating that the holotype presumably 

 originated in the Greater Antilles. We have seen no other specimen, 

 however, with the peculiar basioccipital-presphenoid configuration of 

 M. clinedaphus and conclude that this unique condition is indeed due 

 to distortion. Therefore, we consider M. clinedaphus as the prior name 

 for those bats that up to now have been known as Monophyllus 

 cubanus Miller, 1902. 



Mensural data for the two segments of M. r. clinedaphus are pre- 

 sented in tables 1 and 2. Of the three subspecies of M. redmani, the 

 Cuban and Hispaniolan M. r. clinedaphus is intermediate in size be- 

 tween M. r. redmani of Jamaica and M. r. portoricensis of Puerto Rico. 

 This is demonstrated most clearly by cranial measurements. Fore- 

 arms of M. r. redmani and M. r. clinedaphus are comparable in size ; 

 thus, the forearms are relatively longer in M. r. clinedaphus than in 



