Proceedings of 

 the United States 

 National Museum 



SMITHSONIAN INSTITUTION « WASHINGTON, D.C. 



Volume 124 1968 Number 3649 



The Species of the Subgenus Leptoferonia Casey 

 (Coleoptera: Carabidae: Pterostichus) 



By Hilary A. Hacker * 



The purpose of this paper is to facilitate identification of the 

 species in the subgenus Leptoferonia Casey of the genus Pterostichus 

 Bonelli. As currently defined, the subgenus may be an artificial group- 

 ing of species. After the subgenus Hypherpes Chaudoir has been 

 revised, it may be possible to clarify the relationships between the 

 two subgenera and between the species within Leptojeronia. The 

 subgenus Leptoferonia is composed of 21 species that occur from 

 British Columbia south through California and east to Montana. 

 Seven new species and three new subspecies are described. 



I am very grateful for assistance from a number of people. G. E. 

 Ball, O. L. Cartwright, the K. M. Fenders, and R. L. Hacker reviewed 

 the manuscript and offered many valuable suggestions. O. L. Cart- 

 wright and P. J. Spangler examined the types in the United States 

 National Museum. P. J. Darlington, Jr., lent the types of Pterostichus 

 linearis LeConte and P. longicollis LeConte from the LeConte col- 

 lection in the Museum of Comparative Zoology. K. M. Fender, 

 F. Hasbrouck at Oregon State University, M. H. Hatch at the Uni- 

 versity of Washington, H. B. Leech at the California Academy of 

 Sciences, J. Shuh, and W. C. Stehr at Ohio University also lent 

 specimens. 



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