»o. 3649 LEPTOFERONIA — HACKER 5 



tarsi. In others these setae are absent on all tarsi. One pair of long 

 dorsal setae is always present on the last article (termitiformis has 

 three pairs). There are probably other differences among species in 

 the setae on the legs, but these have not been investigated. The 

 tarsi of the front legs are expanded in the males and narrow in females. 



Sixth abdominal sternum of male. — In the traditional system 

 the last visible sternum, which bears the anal setae, is designated as 

 the sixth sternum. In Leptqferonia all males have a single pair of anal 

 setae, and females have two pairs. In the males of some species, the 

 apical portion of the sternum is thickened. The thickening usually 

 gives rise to a transverse lobe or carina on the ventral surface of the 

 sternum either before the apical marginal impression or at the edge. 

 An unnamed species, which may belong in the subgenus Anillqferonia 

 Van Dyke, has a similar modification of its sixth sternum. In most 

 other subgenera of Pterostichus, carinae, when present, are longitudinal 

 rather than transverse and do not reach the apical edge. 



Male genitalia. — The descriptions of the right paramere refer to 

 the distal portion which is not surrounded by membranes. In sub- 

 genus Hypherpes a short, small, right paramere seems to be the rule; 

 the exact shape varies. In some Leptqferonia and in some other 

 subgenera of Pterostichus the right paramere is elongate. In Hypherpes 

 there is frequently a ridge or lobe on the right side of the ventral 

 surface of the median lobe. This lobe occurs in many Leptoferonia. 

 In some it extends diagonally across the ventral surface. In pumilus 

 the isolated tooth in the middle of the ventral surface appears to be 

 derived from this lobe. In Hypherpes the central area of the distal 

 third of the ventral surface generally is less heavily sclerotized than 

 the adjacent areas. Sometimes the boundary between the left side 

 and tip and the lightly sclerotized area is sharp and there is a clear 

 break between the tip and the shaft on the right side. In some Lepto- 

 feronia the lightly sclerotized area is demarcated clearly and forms 

 a diagonal membranous strip separating the tip and left side from the 

 remainder of the shaft. This condition also occurs in the subgenus 

 Pseudoferonina Ball (1965, p. 107). It can be seen clearly only 

 in material that has been cleared in potassium hydroxide and that 

 is immersed in liquid. Sometimes the tooth in the internal sac of the 

 median lobe is a prominent, dark plate that is curved and tapers to 

 a point at one end, resembling a canine tooth. Sometimes the tooth 

 is small, lightly sclerotized, and obscured by the lips of the gonopore. 

 Sometimes it is absent or replaced by a lobed structure. In the 

 Hypherpes known to me, a large and heavily sclerotized tooth is 

 always present. 



