A REVISION OF THE AMERICAN TWO-WINGED FLIES 

 OF THE PSYCHODID SUBFAMILY BRUCHOMYINAE 



By Charles P. Alexander 



Of the Massachusetts Aijrirultural (^allege, Amherst 



The recent discovery of two undescribed species of the subfamily 

 Briichomyinae, the most generaUzed of the four subfamiHes of Psy- 

 chodidae, makes it advisable to review our knowledge of this group 

 of flies. Tillyard (1926), in writing of the genus Nemopaljms, one of 

 the included genera, remarks as follows: "This genus, placed by 

 some authors in Tanyderidae, unites that family with the Psychodi- 

 dae, and is probably one of the oldest existing types of Diptera." 



I wish to express my deepest thanlcs to the collector, Mr. Raymond 

 C. Shannon, for the opportunity of studying and describing the two 

 new Peruvian species of Bruchomyia. 



Subfamily Bruchomyinae 



1904. Phlebotominae Eaton, Ent. Mo. Mag., ser. 2, vol. 15, p. 55 (part). 



1920. Bruchomyiinae Alexander, Ann. Ent. Soc. America, vol. 13, p. 402. 



1921. Nemopalpinae Edwards, Ann. Mag. Nat. Hist., ser. 9, vol. 7, p. 439. 



1922. Phlebotominae Tonnoir, Ann. Soc. Ent. Belgique, vol. 62>,p. 127 (part). 



1926. Bruchomyiinae Crampton, Ent. News, vol. 37, pp. 33-39, 65-70. 



1927. Bruchomyiinae Alexander, Genera Insectorum, fasc. 189, p. 3. 



1927. Phlebotominae Shannon and del Ponte, Rev. Inst. Bact., Buenos Aires, 



vol. 4, p. 733 (part). 



1928. Bruchomyinae Edwards, Journ. Fed. Malay St. Mus., vol. 14, p. 65. 

 1928. Bruchomyinae Alexander, Proc. Linn. Soc. New South Wales, vol. 53, 



p. 292. 



The use of the name Bruchomyinae, based on the second proposed 

 genus, Bruchoinyia, in preference to Nemopalpinae, erected for the 

 oldest genus, Nemopaljjus, is on the principle of selection of the first 

 proposed group of higher rank than the genus. 



The Bruchomyinae, including only the two genera Nemopalpu» 

 Macquart and Brucliomyia Alexander, appear to represent the most 

 generalized of the recent Diptera. The subfamily is well diflFeren- 

 tiated from the Phlebotominae, with which it has sometimes been 

 placed, by the retention of the primitive arrangement of the branches 

 of Rs, this being dichotomously twice forked. In the Phlebotominae 



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