12 PROCEEDINGS OF THE NATIONAL MUSEUM vol. 122 



specimens of iindescribed salarine genera with vomerine teeth. 

 Hirculops Smith (1959), also with vomerine teeth, is related to Rhabdo- 

 blennius. The combination of characters given in the generic descrip- 

 tion above will serve to separate Eniomacrodus from all other described 

 blennioids. 



Schultz and Chapman (1960) believed Fallacirripectes Schultz and 

 Chapman to be intermediate between Eniomacrodus and Cirripectes 

 Swainson. I believe that Fallacirripectes is the genus most closely 

 related to Eniomacrodus. Fallacirripecies differs from Eniomacrodus 

 in having typically 15, rather than 14, pectoral rays, fewer dorsal 

 spines, fewer segmented dorsal and anal rays, and no vomerine 

 teeth. (Although I have seen no specimens of Sianulus Smith (1959), 

 it appears probable that Fallacirripecies is a junior synonym of 

 Sianulus). 



Intrageneric relationships. — I recognize seven species groups 

 within Eniomacrodus. My recognition of these groups is influenced 

 greatly by the nature (or absence) of the upper lip crenulae and 

 secondarily by other characters. I am unable to delimit intergroup 

 relationships as I do not know which characters are specialized or 

 unspecialized, primitive or advanced. There is no fossil record of 

 the Blenniidae to serve as a guide. 



The species groups I recognize are characterized as follows: 



1. ihalassinus group (E. ihalassinus, E. macrospilus) . Smallest 

 species, lacking crenulae on the upper lip. Indian, western and cen- 

 tral Pacific Oceans. 



2. nigricans group (E. sealei, E. corneliae, E. chiosiictus, E. 

 caudofasciatus , E. nigricans, E. textilis, E. cadenaii, E. vomerinns). 

 Relatively small species, the lateral thirds of the upper lip crenulate 

 and the central portion entire. Circum tropical. 



3. siellifer group (E. stellijer). Relatively large species, the 

 middle third of the upper lip crenulate and the lateral thirds entire. 

 Western Pacific Ocean. 



4. rofeni group {E. rofeni). Relatively small species, the upper 

 lip completely crenulate, high dorsal ray, anal ray and vertebral 

 counts, constantly three pores in the predorsal commissural group, 

 and the dorsal fin free from the caudal. Central Pacific Ocean. 



5. cymaiohiotus group {E. cymaiobioius, E. sirasburgi, E. chap- 

 mani). Relatively small species, the upper lip completely crenulate, 

 generally low meristic counts, and the dorsal fin free from the caudal. 

 Western and central Pacific Ocean. 



6. striatum group (E. striatus, E. jnarmoraius, E. randalli, E. 

 epalzeocheilus, E. niuafoouensis). Relatively large species, the upper 

 lip completely crenulate, generally high meristic counts, and the 

 dorsal fm free from the caudal. 



