178 THE CLASSIFICATION OF THE TILOPTEEIDACE^. 



another very small plant of Scapliospora speciosa was 

 found. In 1896, although again carefully hunted for, no 

 specimens could be obtained, although Haplospora was 

 present in quantity, as on all the previous occasions. 



A fifth visit to the locality was made in April of the 

 present year (1897), with two objects in view : firstly, to 

 conduct, if possible, fertilisation experiments with Tilopteris 

 Mertensii Klitz., which plant occurs in great abundance on 

 the same ground as Haplospora globosa, and secondly to try 

 to get further information as to the relation of Haplo- 

 spora to Scapliospora. The latter object, but not the for- 

 mer, was accomplished. 



In vegetative structure Haplospora glohosa is absolutely 

 identical, histologically, with Scapliospora speciosa^ and 

 Reinke came to the conclusion that these plants were in all 

 probability not only generically, but also specifically, identi- 

 cal (Figs. 1, 2, 3, 4, 5, 6). Kjellman had found antheridia 

 of the Tilopteridean type (Figs. 3 and 7, an.) on a specimen 

 of Haplospora which he described, but did not figure. 

 Subsequently he doubted the accuracj'' of his observation, 

 which Reinke ^ likewise was unable to confirm, although 

 he examined hundreds of specimens of Haplospora. In the 

 course of the present investigation, extending over five 

 years, hundreds, perhaps thousands, of specimens of H. 

 glohosa were examined, but it was not until the .'present 

 year that this question received its final solution through 

 the finding of a specimen bearing well developed, though 

 not mature, sporangia,^ oogoiiia, and antheridia, all per- 

 fectly unmistakable and characteristic (Figs. 7 and 8, s/>., 



1 Cf. Beinke, I.e. Seix Ahdr., p. 2. 



2 The term "sporangium" will be used throughout this paper for 

 the non-sexual reproductive organ of Haplospora, which contains a 

 tetra-nucleate monospore. 



