256 ACCESSOEY MALPHIGHIAN BODY. 



gland in young examples, and sometimes fairly well in old ones, 

 of either sex\ There is generally in each segment a second 

 Malpighian body, which forms the commencement of a tubulus 

 joining that from the primary Malpighian body, and, where the 

 segments are larger, there are three, and possibly in the hinder 

 segments of the Wolffian gland and segments of the kidney 

 proper, more than three Malpighian bodies. 



The accessory Malpighian bodies, or at any rate one of them, 

 appear to have curious relations to the segmental tubes. The 

 necks of some of the anterior segmental tubes (PL xix. fig. 5) 

 close to their openings into the primary Malpighian bodies are 

 provided with a small knob of cells which points towards the 

 preceding segment and is usually connected with it by a fibrous 

 band. This knob is most conspicuous in the male, and in very 

 young animals or almost ripe embryos. In several instances in 

 a ripe male embryo it appeared to me to have a lumen, and to 

 be continued directly forwards into the accessory Malpighian 

 body of the preceding segment. One such case is figured in the 

 middle segment on PL xix. fig. 5. In this embryo segmental 

 tubes were present in the segments immediately succeeding 

 those connected with the vasa efferentia, and at the same time 

 these segments contained ordinary and accessory Malpighian 

 bodies. The segmental tubes of these segments were not, 

 however, connected with the Malpighian body of their proper 

 segment, but instead, turned forwards and entered the segment 

 in front of that to which they properly belonged. I failed 

 to trace them quite definitely to the accessory Malpighian body 

 of the preceding segment, but, in one instance at least, there 

 appeared to me to be present a fibrous connection, which is 

 shewn in the figure already referred to, PL xix. fig. 5, r. sL 

 In any case it can hardly be doubted that this peculiarity of 

 the foremost segmental tubes is related to what would seem 

 to be the normal arrangement in the next few succeeding seg- 

 ments, where each segmental tube is connected with a Mal- 

 pighian body in its own segment, and more or less distinctly 

 with an accessory Malpighian body in the preceding segment. 



* My observations on this subject completely disprove, if it is necessary to 

 do so after Professor Semper's investigations, the statement of Dr Meyer, that 

 segmental tubea in ScylUum open into lymph organs. 



