A NEW SPECIES OF TRICHOSTRONGYLID WORM OF 

 THE GENUS COOPERIA FROM THE CARABAO IN THE 

 PHILIPPINE ISLANDS, WITH A REVIEW OF THE 

 GENUS 



By Benjamin Schwartz 



Senior Zoologist of the Bureau of Animal Industry, United States Department 



of Af/ricuUurc 



Under date of September 9, 1927, Dr. Angel K. Gomez, of the Col- 

 lege of Veterinary Science of the University of the Philippines, 

 forwarded to the writer a portion of the small intestine of a i urabao 

 calf {Buhalus huhalus) from Los Bafios, Laguna, with the informa- 

 tion that the animal had died from inanition and was extremely 

 emaciated before it died. Doctor Gomez said that on post-morten 

 examination the small intestine was found to contain nodules through- 

 out its length and that teased preparations of the nodules revealed 

 the presence of small roundworms. 



Examination of the material by the writer showed the mucosa to 

 be riddled with small, conspicuously raised nodules, varying from 

 about 3 to 5 mm. in diameter, the summit of each nodule being more 

 or less depressed and containing a small opening into a channel of 

 communication between the parasite and the lumen of the intestine. 

 Each nodule contains a single worm which is rather deeply imbedded 

 in the mucosa and is very much twisted, the twists being due, appar- 

 ently, to the technic of fixation. The worms belong to the genus 

 Cooper'ia and represent a heretofore undescribed species for which the 

 name Cooperia nodulosa is proposed. On the evidence this worm 

 must lie regarded as pathogenic and probably of economic importance. 



COOPERIA NODULOSA, new species 



Specific diagnosis — Cooperia: The head (fig. 1) varies in diam- 

 eter with the degree of cuticular expansion and ranges from about 

 47/u. to slightly over 50/^ and bears 4 submedian papillae and 2 amphids 

 or so-called lateral papillae. The esophagus (pi. 1. fig. 1) is slightly 

 less than 0.5 mm. long in the male and from 0.5 mm. to slightly 

 longer in the female, with a maximum diameter of 46/i. The nerve 

 ring is located slightly posterior to the middle of the esoi)hagus. 



No. 2765.— Proceedings U. S. National Museum, Vol. 74, Art. 20 



6044—28 ;j 



