PALEOCENE MAMMALS OF CENTRAL UTAH GAZIN 



37 



have arisen from Protogonodon through Desinatoclaenus. This may 

 well be the case but the larger known forms such as P. pentacus or 

 even P. stenognathus are probably not in the line. It is conceivable 

 that a small form such as P. protogomoides^ whose teeth are closer 

 to Des7natoclaenus than are those of P. pentacus (especially P^), 

 may have given rise to DesmMoclaenus^ assuming a somewhat earlier 

 stage for the Puerco of the San Juan Basin. 



Table 5. — Measurements {in millimeters) of tipper and loioer teeth of 

 Desinatoclaenus hermaeus {TJ. 8. N. M. No. 16202) 



1 Greatest transverse diameter. 



DESMATOCLAENUS PAKACREODUS,so new species 



Type. — Right maxillary portion, U.S.N.M. No. 16201, with M'-M^ 



Horizon and locality. — Wagonroad Paleocene, Dragon Canyon, 

 Emery County, Utah. 



Specifl,c characters. — Larger than Desviatoclaenus hermaeus. Lin- 

 gual portion of upper molars more inflated. M^ relatively larger. 

 Hypocone better developed. 



Description. — A second and somewhat larger species is indicated 

 by material apparently from both the Wagonroad and Dragon hori- 

 zons. The specimen selected as the type. No. 16201 (fig. 20, a), 

 was obtained from the Wagonroad level and includes M^ to M^ The 

 teeth are much like those in Desmatoclaenus hermaeus in most char- 

 acters of the molars, but the lingual portions of these teeth have a 

 more inflated appearance and M=* is relatively larger. Although 

 slightly damaged at the posterointernal angle, M^ shows better evi- 

 dence for a hypocone than in D. hermaeus. The upper molars make 

 an approach toward the conditions seen in Protogonodon stenog- 

 nathus., but the differences, as in Z>. hermaeus., are in the direction of 

 T etraclaenodon. 



A maxillary portion. No. 16177 (fig. 20, &), with M^ and M^ from 

 the Dragon horizon corresponds closely to the type of D. paracreodus, 

 but the teeth being less worn show characters not seen in the type. 

 The external cingulum is weaker than in Protogonodon., and, as in 

 the types of D. hermaeus and D. paracreodus., the cingulum is inter- 

 rupted along the posteroexternal portion of the paracone in M-, and 

 the anteroexternal portion of both teeth projects outward promi- 

 se' TTcpa, near + Kpkai, flesh + 'o«o6s, tooth, in allusion to its resemblance to the carnivore Protogonodon. 



