330 Alexander W . Evans, 



portion is restricted to the upper and outer parts of the sac, the free 

 margin is appressed, and the opening is reduced to a small hole 

 at the outer end. The apex of the lobule, where this hole is situated, 

 never forms a projecting beak as in so many species of Chonanthelia. 

 The water-sac, in fact, bears considerable resemblance to those found 

 in Trachycolea. The lower part of the lobule, usually occupying in 

 F. gibbosa half the length or even less, is the so-called appendiculum 

 of authors. In the present case it is less well developed than in most 

 species of (Iwnanihelia and is variable in form, tending perhaps to be 

 triangular. Along the inner edge the appendiculum is attached to 

 the lobe by a short keel, which spreads more or less widely away from 

 the axis or, in rare cases, is subparallel with it. The outer margin 

 of the appendiculum, extending from the end of the keel to the outer 

 end of the water-sac, is sometimes straight and sometimes more 

 or less strongly curved. In most cases it is well covered over by 

 the lobe but occasionally extends slightly beyond it. The margin 

 is apparently never toothed in any way but may be very slightly cris- 

 pate. In some cases the lobule does not form a water-sac but con- 

 sists of an ovate lamina, which is either plane or slightly canaHc- 

 ulate and which tapers to a blunt point. 



The stylus is remarkably developed and sometimes approaches the 

 lobule itself in size. It varies from an ovate to a broadly reniform 

 expansion and is sometimes plane but much more frequently cris- 

 pate. Sometimes the stylus is appressed to the axis and is directed 

 toward the apex of the shoot (Fig. 9, C, on left) ; sometimes it is com- 

 pletely reflexed by the auricles of the underleaves (Fig. 9, C, on right) ; 

 and sometimes a portion only is reflexed, while the rest occupies the 

 normal position. Under the last conditions the stylus is sometimes 

 bifid. Apparently the broad expansion just described represents 

 an extreme development of the inner basal portion of the stylus. 

 The reason for this idea is that, between the expansion and the lobule, 

 a slender tooth-hke structure, tipped with a hyaline papilla, can be 

 discerned (Fig. 9, B, on right). In most species of FruUania the 

 entire stylus consists of a structure of this character. 



Although the stylus in F. gibbosa is so conspicuous and affords one 

 of the most distinctive features of the species, it is not mentioned 

 by Nees von Esenbeck, and its characteristics are not brought out 

 by Stephani. Gottsche,^ however, emphasizes its importance, and 

 Spruce describes it at some length. According to his account it 

 attains its highest development in the variety densissima, where 



1 Ann. dcs Sc. Nat. Bot. V. 1: i66. 1864. 



