184 BULLETIN 50, UNITED STATES NATIONAL MUSEUM 



B[oiiasa] u[fnbellus] yukonensis Conover, Condor, xxxvii, 1935, 204 in text (crit.), 

 206 (spec, Yukon Territory). — Uttal, Auk, Iviii, 1941, 75, 76 in text (tarsal 

 feathering). 



[Bonasa] [mubellits] yukonensis Todd, Auk, Ivii, 1940, 394 in text. — Uttal, Auk, Iviii, 

 1941, 76, 77, 78 in text (tarsal feathering). 



BONASA UMBELLUS UMBELLOIDES (Douglas) 



Gray Ruffed Grouse 



Adult (gray phase). — Similar to the corresponding phase of Bonasa 

 umbellus umhellus but much less brownish, more grayish and darker, 

 being closest in appearance to the gray phase of phaia from which it 

 differs in being paler gray (and the brown areas paler also) above, less 

 heavily barred below, and with a longer part of the tarsus feathered. 

 The gray of the feathers of the nape, back, rump, upper tail coverts, 

 and the tail smoke gray lightly vermiculated with fuscous to blackish ; 

 the brown areas of the interscapulars cinnamon-brown to russet heavily 

 blotched with black; the brown of the upper surface of the wings dull 

 Saccardo's umber speckled and washed with grayish ; below more heavily 

 banded than umbellus, less so than phaia, the brown bands averaging 

 slightly darker than in umhellus — pale tawny-olive darkening on the 

 sides and flanks to sepia ; tarsus feathered for more than half its length. 



Adult (brown phase). — Similar to the gray phase but with the tail 

 tawny-olive instead of gray and interscapulars and upper surface of the 

 wings slightly more extensively brownish, and the breast and upper ab- 

 domen averaging more washed with tawny-buff. 



Juvenal (brown phase). — Similar to the corresponding plumage of 

 B. u. umhellus but less rufescent, more grayish, the general color of the 

 upperparts of the head, body, and wings being buffy brown to grayish 

 olive-brown, the rectrices between wood brown and drab ; ventral barring 

 darker — grayish buffy brown. 



Juvenal (gray phase). — Like the brown phase but gray — the ground 

 color of the upperparts of the body and wings and tail being grayish drab 

 to ashy hair brown ; ventral barrings darker — mummy brown, the under- 

 parts less washed with buffy. 



Downy young. — Not distinguishable from that of the nominate race. 



Adult male.—Wmg 171-195 (182.6); tail 144-174 (157.7); culmen 

 from base 24.4-29.4 (26.3) ; tarsus 39-45.5 (42.5) ; middle toe without 

 claw 35.0-40.6 (37.5) ; unfeathered part of tarsus 10.9-24.0 (17.7) mm.).^^ 



Adult female— Wing 169-180 (1746) ; tail 125-134 (130.4) ; culmen 

 from base 23.6-27.6 (25.6) ; tarsus 37.5-43.2 (41.0) ; middle toe without 

 claw 33.8-36.3 (349) ; unfeathered part of tarsus 10.9^17.8 (15.3 mm.).so 



'* Forty-three specimens from Alberta, northern British Columbia, and Montana. 

 "° Eleven specimens from Alberta, British Columbia, Montana, and northwestern 

 Wyoming. 



