MILLIPEDS — HOFFMAN 217 



first the progenitors of the present crassicutis phratry and then the 

 group now recognized as Epeloria. Of the existing pachydesmids, 

 incursus and duplex have gonopods most like those of clarus, and are 

 in fact very similar to each other. They occupy respectively the 

 northwestern and northeastern corners of the generic range, suggest- 

 ing slightly modified remnants of their ancestral stock. Near the 

 center of the range, a population reinvaded the southern end of the 

 Appalachians along the old north-south course of the Tennessee 

 River. This population evolved a greater size and more complex 

 gonopod structure, and now, as the laticollis phratry, separates the 

 eastern and western subspecies of the older crassicutis phratry. 



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