ART. TWO-WIXGED FLIES OF TRIBE MILTOGRAMMINI ALLEN 85 



inens of HilareUa, and decerifi, the typo of Enmacr onychia, I am con- 

 vincod that the hitter is generically distinct. The species of Enma- 

 cronycJiia as a whole possess ahnost the same comhination of charac- 

 ters as those of Gy7nno2)7'osopa, and certain species such as elita and 

 rohweri possess striking resemblances to both genera, but the male 

 genitalia in EumacronycMa are uniformity large and highly specialized, 

 somewhat resembling those of typical Sarcopliagiiiae, while the male 

 genitalia of Gymnoprosopa are small and relatively unspecialized as in 

 Metopia and related genera. In Eumacronychia, the female genital 

 segments extrude conspicuously, the apical scutellar bristles are 

 absent or if present are much smaller than either pair of lateral 

 marginals, the front is usually distmctly narrower near base of the 

 antennae than towards the vertex, and the male usually possesses 

 conspicuously long broad pulvilli, and a tendency toward villosity on 

 the legs, all characters distinguishing it from Gyinnoprosopa. Other 

 characters of the genus are as folknvs: 



Inner orbits not sinuous or pinched in at the level of vibrissae; 

 frontal vitta light red or yellow, at least as wide as the parafrontal, 

 diverging moderately toward the vertex; iunule scarcely visible; a 

 single frontal row of bristles terminating at base of antennae, not 

 suddenl}?- divergent below; proclinate ocellar bristles present; orbitals 

 in both sexes; antennae distinctly more than half as long as the face; 

 penultimate joint of the arista short; vibrissae not more than the 

 length of second antennal joint above front edge of oral margin; 

 in profile, head length at vibrissae sometimes subequai, usually much 

 less than at base of antennae; lower part of bucca with sparse black 

 bristly hairs; no pale hairs about the oral cavity; proboscis moderately 

 stout, shorter than the head height, with fleshy labella and normal 

 palpi. Thorax with three strong postsutural dorsocentral bristles 

 and two sternopleurals. Pollen on the intermediate segments of the 

 abdomen arranged in basal bands which are not scalloped or indented 

 on the apical margin; abdomen never marked with large black spots. 

 Wing with apical cell open; only the third vein bristly; last section 

 of fifth vein less than half the length of the preceding section. Joints 

 of the fore tarsus not ornamented, conspicuously reduced in size, 

 flattened, or otherwise modified from the normal in either sex; pul- 

 villi of male variable in length. 



None of the species of this genus are known at present from outside 

 the continental North American area. I have seen very few speci- 

 mens from north of the latitude of Nc w Jersey. The genus is com- 

 posed of several species of attenuated or possibly discontinuous dis- 

 tribution. Not one of the species is definitely known to have a 

 transcontinental range except sternalis identified from Galveston, 

 Texas, and Granada, Nicaragua. The Southv/est appears to be the 

 center of distribution of the jjroup. Nothine: is known concernincr 



