ART. 4 BEETLES OF THE GENUS COCCINELLA DOBZHANSKY 



that used by other authors. The part functioning in Coccinellidae 

 as a penis is, as shown by its development, homologous to only the 

 proximal part of the penis of other Coleoptera (according to the 

 unpublished data of the author). This part is termed the sipho 

 (penis of other authors). Moreover, the penis, which is homologous 

 to the distal part of the penis of other Coleoptera, is intimately fused 

 with the basal plates (basal piece of other authors). The fingerlike 

 paramera (lateral lobes) are articulated with the basal plates. The 

 trabes (tegminal strut of other authors) is an unpaired chitinous rod 

 articulated to the basal plates, and connected by muscles with the 

 proximal end of the sipho. 



In the genus Coccinella the sipho (s, fig. 1 and figs. 13-20) is hook- 

 shaped. Its proximal end, in 

 most species, is strongly chiti- 

 nized and separated from the 

 body of the sipho to form the 

 so-called siphonal capsule {sc, 

 figs, land 13-20). Onlyin(7oc- 

 cinella undecimpunctata Lin- 

 naeus the siphonal capsule is 

 rudimentary (fig. 20). The 

 distal end of the sipho carries bS-.. 

 rather strongly developed 

 praeputial sacs. The penis 

 {p, figs. 1, 2-12) frequently 

 possesses complicated proc- 03— 

 esses on its distal end. The 

 form of the penis is exceed- 

 ingly variable and constitutes 

 the best specific character. 

 The trabes {tr, fig. 1) is rela- 

 tively short and thick, and its 

 free end has no clearly pro- 

 nounced emargination. The basal plates {bp, fig. 1) are strongly 

 developed. Paramera (pa, figs. 1, 2) are fingerlike and slightly 

 compressed from the sides. 



Female genitalia. — The receptaculum seminis is large, mostly rather 

 clearly differentiated into the cornu (c, figs. 22, 28), the nodulus {n), 

 and the ramus (r). The sculpture of the walls of the receptaculum, 

 consisting of chitinous rings, is well developed in all species except 

 Coccinella undecimpunctata Linnaeus and its relatives. The infundib- 

 ulum (?', figs. 22, 28) has a funnel-shaped dilatation on its anterior end 

 and, in some species, a similar dilatation on its posterior end. The 

 ductus receptaculi (dr, fig. 28) is very short and nearly hidden in the 



Figure 1. — Male genitalia of Coccinella novemnotata 

 Herbst (lateral view), bp. Basal plates; p, penis; pa, 

 paramera; s, sipho; sc, siphonal capsule; tr, trabes 



