FROGS OF COLOMBIA — COCHRAN AND GOIN 89 



Genus Pseudis Wagler 



1830. Pseudis Wagler, p. 203 (type species, Rana paradoxa Linnaeus). 



Generic diagnosis. — Pupil horizontal. Tongue circular, entire and 

 slightly free behind. Vomerine teeth present. Tympanum distinct. 

 Fingers free, first finger opposable to the others; toes completely 

 webbed; tips of all digits pointed or swollen into small disks. Outer 

 metatarsals separated by a web. Omosternum cartilaginous; sternum 

 a cartilaginous plate. Terminal phalanges simple. 



Pseudis paradoxa nicefori, new subspecies 



Plates 12g, 68a-c 



Holotype. — MLS 443, an adult from Monteria, Cdrdoba, Colombia, 

 collected by Hno. Niceforo Maria, February 22, 1966. 



Paratopes. — USNM 145076 (larva), Rio San Jorge, between 

 Lagunas Puerco and Sabanas, Bolivar; USNM 145780 (larva), 

 Rioviejo, lower Rio Magdalena, Bolivar; CNHM (larva), V&lledupar, 

 Badillo, upper Rio C6sar, Magdalena. 



Diagnosis. — Vomerine teeth in two large, transverse, well-separated 

 series between the choanae; greatest diameter of tympanum equal to 

 that of eye and equal to distance from eye to nostril; a thick glandular 

 ridge from posterior corner of eye, continuing above tympanum and 

 shoulder to fade out behind the level of the axilla; lower surface of 

 femur with heavy gray reticulations enclosing pale round or oval 

 and sometimes irregular areas, with a wide pale stripe along anterior 

 surface of femur above the reticulations; lower surface of tibia pale 

 with a few thin gray vermiculations; venter olive-buff with some 

 small gray spots on throat, more definitely aligned along center of 

 chest; a large triangular light spot outlined with gray below shoulder 

 attachment; belly immaculate; back with two very irregular longi- 

 tudinal rows of darker spots on each side (one row near the midline, 

 the other in the dorsolateral area) ; a large light transverse spot 

 immediately behind anus and a second smaller patch behind it. 



Description of holotype. — Vomerine teeth in two very heavy, 

 transverse, well-separated series between the choanae; tongue two- 

 thirds as wide as mouth opening, deltoid, attached except on its 

 extreme border, which is straight; snout moderately short, rounded 

 when viewed from above, rounded in profile, the upper jaw extending 

 well beyond the lower. Nostrils superior, slightly projecting, their 

 distance from end of snout almost as great as their distance from eye. 

 Can thus rostralis indistinct; loreal region barely concave, sloping 

 broadly to the upper lip. Eye moderate in size, prominent, its diameter 

 two-thirds its distance from tip of snout; interorbital diameter twice 

 that of upper eyelid, equal to interval between nostrils. Tympanum 



