36 U.S. NATIONAL MUSEUM BULLETIN 288 



Bibron, 1841, p. 638— Giinther, 1858, p. 90.— Dunn, 1944c, p. 491; 



1957, p. 77.— Cochran, 1961, p. 107, fig. 39.— Gorham, 1963, p. 25. 

 1847. Phyllobates melanorrhinus Berthold, p. 14, pi. 1, fig. 7 (type locality, 



Popayan, [Cauca,] Colombia). 

 1923. Phyllobates nicefori Noble, p. 1 (type locality, "town of Andes," 



[Antioquia,] Colombia). 



Description. — USNM 137360, an adult female from Popay&n, 

 Cauca, Colombia. Head rather short; tongue half as wide as mouth 

 opening, ovate, its posterior border free and unnotched; snout broadly 

 rounded when viewed from above, truncate in profile, the upper 

 jaw extending well beyond the lower. Nostrils lateral, projecting, 

 their distance from end of snout one-half their distance from eye. 

 Canthus rostralis rounded; loreal region flat, vertical, in line with 

 the vertical upper lip. Eye large, prominent, its diameter equal to 

 its distance from tip of snout; interorbital diameter one-half that of 

 upper eyelid, equal to interval between nostrils. Tympanum very 

 distinct anteriorly, less so posteriorly, its greatest diameter two- 

 thirds that of eye, separated from eye by an interval equal to one- 

 fourth its own diameter. Fingers moderate in length, without lateral 

 ridges, free, with distinct disks; first finger a little longer than second 

 and equal to fourth, the tips of first and fourth reaching to base of 

 penultimate phalanx of third, the disk of which covers about one-fourth 

 the tympanum; a small round thumb pad present; a larger palmar 

 callus; metacarpal tubercles well developed. Toes free, with faint 

 lateral ridges, third toe longer than fifth, its disk reaching halfway 

 on antepenultimate phalanx of fourth; disk of fourth toe covering 

 one- third the tympanum and slightly larger than that of third finger; a 

 small oval inner metatarsal tubercle and a still-smaller, round outer 

 one; a slight tarsal ridge ending in a heavy diagonal gland midway 

 to heel ; a heavy skinfold on heel and knee. Body moderately elongate, 

 in postaxillary region narrower than greatest width of head. When 

 hind leg is adpressed, heel reaches center of eye; when limbs are 

 laid along the sides, knee and elbow overlap; when hind legs are 

 bent at right angles to body, heels touch. Skin of upper parts smooth 

 (finely pustular under the lens) and very resistant to abrasion; venter 

 smooth, except for a few small granules on posterior part of abdomen 

 and on lower femur beside the vent; no glandular ridge from posterior 

 corner of eye; a very slight skinfold across chest; no ventral disk; 

 no external vocal sacs apparent in the male. 



Dimensions. — Head and body, 42.5 mm.; head length, 14 mm.; 

 head width, 13.5 mm.; femur, 18.5 mm.; tibia, 20 mm.; foot, 16 mm.; 

 hand, 12 mm. 



Color in alcohol. — Dorsum uniform pearl gray; upper limb surfaces 

 with numerous, very small slate-gray dots which are set closer to- 



