28 U.S. NATIONAL MUSEUM BULLETIN 2 88 



black; a large white spot on loreal region extending below eye and 

 another on tip of snout. (In life, the silvery- white spots were rich 

 geranium-pink, as traces of that color still remain on some other 

 individuals of this lot.) 



Remarks. — The anomaly of possibly three subspecies of a single 

 species occurring in the Choc6 can be explained by plotting their 

 distribution on a map. One form, D. tinctorius histrionicus, comes from 

 inland and southward, in the valley between the Central and the 

 Western Cordilleras. Another, D. t. chocoensis, appears to be only from 

 the vicinity of Andagoya. The third one, D. t. wittei, is found from the 

 seacoast west of the Western Cordilleras, along the Rio Pepe to 

 Andagoya and somewhat to the east. Near Andagoya are watersheds 

 for three rivers — the Atrato, running north to empty into the Carib- 

 bean, the San Juan, running south and west to the Pacific; and the 

 Baud6, flowing west, more directly to the Pacific. Specimens from 

 Sierra de Baud6 have been classified as probable intergrades between 

 D. t. wittei and D. t. chocoensis, but intensive collecting is needed to 

 decide the advisability of putting wittei into the synonymy of the latter. 



Specimens Examined 

 COLOMBIA 



Choc6: One km. west of Andagoya, USNM 124218-32; Andagoya, BM 

 1915.10.21.58-61, 1915.10.21.63-5, USNM 145791; Bahfa de Utria, USNM 

 144991; Boca de la Raspadura, Rio San Pablo, AMNH 13711-2, 13716-9, 

 USNM 147510; Finca La Victoria, near Pepe" on shore of Rio Baud6, USNM 

 144986-90; Upper Condoto, 12,000 ft., BM 1913.11.12.90-3; upper Rio 

 Quito, upper bank of Rio San Pablo, AMNH 13695-701, 13703-5, 13707-9, 

 USNM 147508-9. [Probable intergrades: wittei X chocoensis, Sierra de Baud6, 

 ANSP 25617-25.] 



Dendrobates tinctorius histrionicus Berthold 



Plate 4g-i 



1845. Dendrobates histrionicus Berthold, p. 43, pi. 1, fig. 8 (type locality, New 

 Granada).— Funkhouser, 1956, p. 76, 3 figs.— Gorham, 1963, p. 25. 



1864. Dendrobates tinctorius cocteani (lapsus, for cocteaui) Steindachner, p. 261 

 (type localities, Brazil and Colombia). 



1913. Dendrobates tinctorius cocteaui. — Boulenger, 1913, p. 1027, pi. 104, figs. 2-4. 



1942. Dendrobates tinctorius histrionicus. — Laurent, 1942, p. 12, figs. 



1944. Dendrobates histrionica. — Dunn, 1944c, p. 520. 



Description. — USNM 139778, an adult from 75 miles south of 

 Andagoya, Choc6, Colombia. Tongue almost half as wide as mouth 

 opening, oval, its posterior border free and unnotched; snout truncate 

 when viewed from above, flat and sloping backwards in profile, the 

 upper jaw extending well beyond the lower. Nostrils lateral, slightly 

 projecting, their distance from end of snout about two-fifths their 

 distance from eye. Can thus rostralis distinct; loreal region flat, vertical, 



