FROGS OF COLOMBIA — COCHRAN AND GOIN 133 



throat, and elongate olive spots running longitudinally on belly; a 

 sepia lateral line beginning at tip of snout and running along the 

 canthus, enlarged on top of head to the dark spots bordering the 

 median light line of snout, continuing and widening greatly behind 

 eye to groin; upper lip cream buff with two wood brown spots on its 

 margin, below and in front of eye; palms of hands and soles of feet 

 cream buff, with wood brown spots across penultimate phalanges 

 of the three outer toes and fingers; a large brown patch on outer 

 half of tarsus and similar but paler markings on lower surfaces of 

 tibia and femur. 



Remarks. — The above-described example is nearly identical with 

 Boulenger's figure of his type specimen from Pefia Lisa, Choc6, which 

 was only a half-millimeter longer. Dimensions (in mm.) of a young 

 frog (MLS 172, from Manzanares, Caldas) are as follows: head and 

 body length, 19.5; head length, 6.5; head width, 6.5; femur, 9; tibia, 

 9.5; foot, 7; hand, 5.5. The light lines on its back are finer and there 

 are fewer spots on its belly; otherwise it closely resembles the Puerto 

 Utria example. Ten specimens from El Valle, Chocd, agree well with 

 the first two mentioned, except that two of them have nearly im- 

 maculate bellies. The adpressed heel reaches between the anterior 

 corner of the eye and the nostril in all except one example, in which 

 it reaches only to the center of the eye. 



The femur averages a greater proportional length than in any other 

 Colombian atelopid, although it is approached by A. varius elegans 

 and A. v. glyphus in this characteristic. The tibia is also very long in 

 all three forms. 



The probable relationship of this species to A. varius bibroni is 

 considered in the discussion under the latter. The characters by which 

 Barbour separated his subspecies A. spurrelli certus from nominate 

 A. spurrelli — the breaking up of the dark dorsal areas of spots, the 

 finer spotting of the belly in males, and the nearly immaculate belly 

 in females of certus — are subject to much variation in examples from 

 both Panama and Colombia. For the present, therefore, it seems best 

 to retain the name spurrelli for all of them. 



Specimens Examined 

 COLOMBIA 



Caldas: Manzanares, MLS 172. 



Choc6: El Valle, USNM 151280-9; Puerto Utria, USNM 110958; Bahia 

 Solono, USNM 144933-5. 



Atelopus varius elegans (Boulenger) 



Plate 19d-f 



1882. Phryniscus elegans Boulenger, 1882c, p. 464 (type locality, Tanti, Ecuador, 

 2,000 ft.); 1882a, p. 155; 1891b, p. 134, text fig.— Nieden, 1926, p. 81. 

 337-262—70 10 



