264 U.S. NATIONAL MUSEUM BULLETIN 2 88 



red and variabilis has one that is flesh-colored to yellowish. We have 

 never seen a live specimen of bogerti, so we do not know the color of 

 its patagium, but in preservative it does not seem to be heavily 

 pigmented. 



In a way, this chain of species seems to be the ecological equivalent 

 of the three forms (H. labialis krausi, H. I. labialis, and H. I. platy- 

 dactyla) with brightly colored patagia that range from south to north 

 in the Oriental Andes of Colombia. Of these we have seen only H. I. 

 labialis alive; its patagium is bright blue in life rather than reddish or 

 yellowish as seen in columbiana and variabilis. While both groups do 

 have well-developed patagia and seem to be ecological equivalents, 

 one of the other, we do not imply that they are closely related, for we 

 are not at all sure this is the case. 



We take pleasure in naming this species for Dr. Charles M. Bogert, 

 whose kindness in lending material has contributed so much to the 

 present work. 



Hyla columbiana Boettger 



Plate 35a-c 



1892. Hyla columbiana Boettger, p. 41 (type locality, Colombia, Cauca, Pop- 

 ayan).— Mertens, 1922, p. 164.— Nieden, 1923, p. 265.— Rivero, 1964a, 

 p. 303. 



1944. Hyla colombiana. — Dunn, 1944a, p. 77. 



Diagnosis. — A small frog without a distinct pattern on the back, 

 with a well-developed patagium, and with reduced webbing between 

 the fingers. 



Hyla columbiana is most closely allied to variabilis and bogerti. It 

 differs from the former in lacking extensive mottling on the chin, 

 throat, and belly, and from the latter in having a longer leg (the ex- 

 tended heel reaching the anterior rather than posterior margin of eye) 

 and in lacking a well-developed pattern on the back. Preserved speci- 

 mens sometimes closely resemble H. minuta but differ in lacking both 

 the distinct pattern on the back and the transverse white line above 

 the vent and across the base of the heels. 



Description. — MLS 192b, Popayan, Cauca, Colombia. Vomerine 

 teeth in two short, transverse series, lying between the small, rounded 

 choanae; tongue one-half as wide as mouth opening, rounded, its 

 posterior border slightly free and unnotched. Snout short, rounded 

 when viewed from above, rounded in profile, the upper jaw extending 

 a little beyond lower; nostrils as lateral as superior, not projecting, 

 their distance from end of snout about one-half that from eye, sep- 

 arated from each other by an interval equal to about their distance 

 from eye. Canthus rostralis weakly defined; loreal region nearly flat 

 and nearly vertical, the upper lip not flaring out below it. Eye mod- 

 erate, not particularly prominent, its diameter greater than its dis- 



