362 U.S. NATIONAL MUSEUM BULLETIN 2 88 



its own greatest diameter. Tympanum lying just below distal end 

 of a postorbital bony ridge. Fingers slightly webbed at base, fourth 

 finger slightly shorter than second, just reaching to base of penul- 

 timate phalanx of third; a slightly projecting rudiment of a pollex; 

 no ulnar ridge (broken right radio-ulna shows in X-ray); toes free 

 of web, third toe a phalanx longer than fifth; a distinct oval inner 

 and a smaller but distinct rounded outer metatarsal tubercle; no tarsal 

 ridge; no dermal appendage on heel. Body short, in postaxillary 

 region much narrower than the greatest width of head; when hind 

 leg is adpressed, heel reaches to eye; when limbs are laid along the 

 side, knee and elbow slightly overlap; when hind legs are bent at 

 right angles to body, heels fail to meet by several millimeters. No 

 patagium or axillary folds from the back of upper arm to side of 

 body. Skin of upper parts dry and slightly granular. Top of head with 

 parallel longitudinal bony ridges; a prominent bony postorbital 

 ridge on each side. Skin along the side below sacral hump somewhat 

 wrinkled. A rather narrow glandular ridge passing down from the 

 tip of the postorbital ridge and behind tympanum; skin of throat 

 and chest smooth but with scattered tiny wart-like tubercles; that 

 of belly and lower surface of thigh uniformly and rather coarsely 

 granular; no traces of a skinfold across chest; no inguinal gland; 

 adult female, no vocal sac apparent. Skin of head not co-ossified with 

 skull, roof of skull heavily exostosed. 



Dimensions. — Head and body, 51.5 mm.; head length, 21.4 mm.; 

 head width, 28.4 mm.; femur, 22.7 mm.; tibia, 21.3 mm.; heel-to-toe, 

 34 mm.; hand, 18.2 mm. 



Color in alcohol. — Uniform pale brown, slightly darker along the 

 side between axilla and groin. 



Remarks. — In their presentation on the genus Amblyphrynus , 

 Cochran and Goin (1961, pp. 545-546) remarked as follows: 



We are not at all sure of the exact relationships of Amblyphrynus. The very 

 broad head gives it an aspect similar to Ceratophrys or Lepidobatrachus but we 

 are not at all convinced that the broad-headed leptodactylids form a natural 

 group. At any rate, Amblyphrynus differs from Ceratophrys in lacking toe webs 

 and supraciliary processes, in having the cranial derm free of the head, and in 

 having the vomerine teeth well behind the choanae. From Lepidobatrachus it 

 differs in having the toes free of webs, in lacking paired odontoids at the mandib- 

 ular symphysis, in having the tongue free behind, and in having the cranial derm 

 free of the head. 



If, on the other hand, toe webbing is indicative of relationship in the lepto- 

 dactylids, then on the basis of free toes and having the terminal phalanx of the 

 digits pointed and stylus-like rather than T-shaped, Amblyphrynus belongs with 

 the group that includes Eusophus, Zachaenus, and perhaps Craspedoglossa, al- 

 though Craspedoglossa is somewhat intermediate and has the tip of the terminal 

 phalanx with a round knob rather than being either truly pointed or T-shaped. 

 From Eusophus, Amblyphrynus differs in having the vomerine teeth well behind 



