364 U.S. NATIONAL MUSEUM BULLETIN 2 88 



being 18 mm. in length) indicates that it may be an immature frog, 

 but its true status must remain in doubt until more collections have 

 been made at the type locality. 



The characteristics of Borborocoetes columbianus are given by 

 Werner and by Nieden. The species is said to be near B. quizensis 

 Jiminez de la Espada, from which it differs in the following points: 

 Tympanum present, more or less distinct, two-thirds the diameter of 

 eye; vomerine teeth behind the choanae; first finger as long as second; 

 heel reaching to anterior margin of eye; nostrils much closer to tip 

 of snout than they are to eye; subarticular tubercles distinct but 

 small; inner metatarsal tubercle very small, no outer one; skin smooth. 



Above blackish brown; a light band between the eyes; posterior 

 thigh with fine dark dots; belly white with brown vermiculations ; 

 thigh below reddish brown. 



Genus Ceratophrys Wied 



1824. Ceratophrys Wied, 1824b, column 672 (type species, C. varius Wied). 



Generic diagnosis. — Pupil horizontal. Tongue heart-shaped, free 

 behind. Vomerine teeth present. Tympanum usually distinct, some- 

 times hidden. Fingers free; toes somewhat webbed, the tips not 

 dilated. Outer metatarsals united. Omosternum cartilaginous; sternum 

 a cartilaginous plate which ossifies in the fully adult state. Terminal 

 phalanges simple. Upper eyelid often produced into a somewhat 

 distinct horn or series of warts. 



The two species that occur in Colombia can be quite readily dis- 

 tinguished from each other by their differences in leg length and other 

 characters. 



Key to Colombian Species of Ceratophrys 



a 1 . When hind leg is adpressed, metatarsal tubercles reach to between tympanum 

 and eye, and heel reaches barely to axilla; size to 86 mm . . C. calcarata 



a 2 . When hind leg is adpressed, metatarsal tubercles reach beyond eye, and heel 

 reaches at least to angle of mouth; size to 106 mm C. cornuta 



Ceratophrys calcarata Boulenger 



Plate 45d-f 



1890. Ceratophrys calcarata Boulenger, p. 327, pi. 26 (type locality, Colombia).- — 

 Boettger, 1892, p. 29.— Ruthven, 1922, p. 53.— Nieden, 1923, p. 384.— 

 A. Lutz, 1927, p. 40.— Dunn, 1944c, p. 509.— Gines, 1959, p. 110, 

 fig.' — Cochran, 1961a, photograph p. 89. — Rivero, 1961, p. 83. — Gorham, 

 1963, p. 16. 



Description. — USNM 117510, a young female from Rio Guaimaral, 

 Atlantico, Colombia. A thick crest along can thus, along inner border of 

 eye, and above tympanum ; vomerine teeth in two short ridges, border- 



