FROGS OF COLOMBIA — COCHRAN AND GOIN 371 



given by Dunn (1949, map, p. 8), and the two frogs from Santa Rita, 

 Solimoens River, Brazil, constitute a record for western Brazil. 

 Persistent search should bring to light more specimens from the 

 Amazon and its tributaries, perhaps even from the Putumayo drain- 

 age, as Dunn suggested. 



Until more of these rare frogs have been collected and studied, 

 it seems best to consider as full species Boulenger's Edalorhina 

 buckleyi, 1882, and his E. nasuta, 1912, and Cope's Bubonius plicifrons, 

 1874. 



Specimens Examined 

 COLOMBIA 



Amazonas; Leticia, JRT 104; Puerto Narino, USNM 146954-6. 

 BRAZIL: Amazonas: Santa Rita, Solimoens River, BM 1915.3.9.16-7. 

 ECUADOR: Canelos, USNM 145790, BM 80.12.5.194a-c; Puerto del Mairo, 



BM 81.7.8.11; Sarayacu, BM 80.12.5.283. 

 PERU: Huanuco: Ceja de Montana, JP 0747 (7). Orellana: Santa Clara, 



Sta. G5, USNM 127182; Domo Santa Clara, USNM 127928-31; Parene, 



BM 1900.11.27.45. Satipo: Marankiari, JP 0058. 



Genus Eleutherodactylus 



1841. Eleutherodactylus Dumeril and Bibron, p. 620 (type species, E. 



martinicensis) . 



Generic diagnosis.- — Pupil horizontal. Tongue subcircular or oval, 

 entire or slightly notched and free behind. Vomerine teeth present. 

 Tympanum generally distinct. Fingers free or nearly so; toes free or 

 webbed; all digits with dilated disks. Outer metatarsals united. 

 Omosternum cartilaginous; sternum a cartilaginous plate. Terminal 

 phalanges T-shaped. 



The members of the genus Eleutherodactylus have received at 

 least 20 generic names. Myers (1962, pp. 195-302) has made an 

 appraisal of all these names and his work should be consulted as an 

 aid in the study of this genus. 



Eleutherodactylus is very diversified in northwestern South America, 

 where speciation is high and variation within a species in nearly all 

 characters is considerable; hence, the constructing of an adequate 

 "key" in the style of Boulenger and Nieden has become almost impos- 

 sible. 



The members of this genus appear to fall into natural groups, how- 

 ever, and it is by defining the predominant characters of each group 

 that the following key has been constructed. The student not already 

 familiar with this puzzling genus may find it necessary in keying a cer- 

 tain frog to compare several descriptions of those forms lying nearest 

 to that selected in the key, as such features as forward extension of heel, 

 degree of webbing, development of disks, and other characters vary 



