386 U.S. NATIONAL MUSEUM BULLETIN 288 



siderably longer than the second in the remaining 75 percent. The dor- 

 sal skin is faintly to heavily granular in more than half the examples, 

 pustular or shagreened in the others. 



The dark postanal patch is distinct in less than half the examples at 

 hand. There are two or three black shoulder tubercles in nearly all 

 specimens. The knee has wide dark diagonal stripes near it in nearly all 

 specimens. The venter is spotted in about 66 percent of the specimens 

 and is immaculate in the remaining 34 percent. The throat has a 

 median light stripe down its center in 75 percent. 



While there is no significant variation in the six critical measure- 

 ments of E. raniformis (74 examples from Antioquia to Narifio) and 

 of E. longirostris, including palmatus and Dunn's "Jitzingeri" (52 

 examples from Nicaragua, Costa Rica, Panama, Ecuador, and Chocd 

 to Putumayo in Colombia), there nevertheless seems to be a fairly 

 constant color difference. The throat in longirostris usually is some- 

 what to quite dark-spotted, but in many Panamanian examples 

 there is a white median area down the center of the throat, with the 

 dark spots grouped in a more-or-less regular longitudinal border on 

 either side of it. This same throat pattern occurs in CNHM 54384-90 

 and 54588 from Caldas. The throat of raniformis is immaculate. Often 

 there are three black tubercles marking the shoulder in longirostris, 

 while one (or sometimes two at the most) occurs there in raniformis. 

 A distinct dark triangular patch covering the postanal area is found 

 in many examples of longirostris, this patch being especially evident 

 in the young. No definite postanal marking can be seen in raniformis, 

 although the postanal region is often suffused with a dark tone. 



Some of the Valle examples, freshly collected, are somewhat like 

 the Panama frogs identified by Dunn as fitzingeri except that the 

 former have a slightly longer snout, a smaller tympanum, and much 

 smaller light dots on the posterior femur. 



Most young examples show more of the fine granulations on the 

 belly than do the adults. The pattern is often more distinct, consisting 

 of two wide dark chevrons, the first above the shoulders, the second 

 just before the sacrum, and continuing on the sides as diagonal 

 stripes. The throat is often pale in the center, and with dark spots 

 on both sides. The very dark distal bar on the femur and the dark 

 spots on the knee are usually present, and are diagnostic of this 

 species. 



Specimens Examined 

 COLOMBIA 



Antioquia: Jeric6, AMNH 22605-7, Chigorod6, USNM 151892; Ventanas, 



50 km. northwest of Yarumal, MLS 353. 

 Caldas: Pereira, MLS 317. Near Salamina, north of Manis, MLS 321; Santa 

 Cecilia, Pueblo Rico, CNHM 54384-90, 54588-9 ;Villa Maria, USNM 146974-9. 

 Cauca: Gorgona Island: Prison Camp, USNM 145137. 



