328 U.S. NATIONAL MUSEUM BULLETIN 288 



terminates above the vent. In the broadened portion of the dark band 

 above the urostyle there is a small, transverse, dumb-bell shaped 

 patch of the lighter dorsal ground color. The thigh, leg, and foot are 

 each crossed by two broad bands of dark brown which are coincident 

 when the legs are flexed. A similar dark brown band crosses each 

 forearm just above the wrist. A stripe of dark brown originates behind 

 each eye and passes posteriorly around (but not through) the tym- 

 panum and terminates above the insertion of arm. The ventral ground 

 color is about the same tone of brown as the dorsal ground color. 



Variation. — The problem of variation in this species is no nearer 

 solution today than it was when Duellman (1956) published his study. 

 (For a study of the variation outside of Colombia, we refer the reader 

 to Duellman's report.) None of the type localities of the six separate 

 names presently listed as synonyms are within Colombia. In addition 

 to these six available names, there are several other forms now mas- 

 querading as Hyla which, we suspect, will be referred to the genus 

 Phrynohyas when they have been studied. 



We note three types of variants in the Colombian material at hand. 

 Two specimens from Astillero and one specimen from Leticia have 

 conspicuous dorsal patterns of dark brown blotches superimposed 

 on a lighter ground color, together with distinct bands across the 

 thighs, shanks, and feet. These apparently are what Dumeril and 

 Bibron called Hyla vermiculata. Four other specimens, all from La 

 Providencia, Caqueta, seem to represent the variant described by 

 Spix as H. bujonia. These four specimens are not only smaller (60.3, 

 56.0, 44.6, and 42.8 mm. in head-body length) but are also more 

 slender, elegant-looking frogs. 



The remaining ten specimens from Colombia now before us are stout- 

 bodied, more or less uniformly brownish frogs. Some have indistinct 

 dark crossbands across the legs, others show no evidence of such 

 markings. A few have faint, small spots on the dorsum. Goin and Layne 

 (1958, p. 112) pointed out that there is a distinct difference in the 

 number of maxillary teeth between specimens from Leticia, in Colom- 

 bia, and from Trinidad. The number of maxillary teeth on the left side 

 of our four specimens from Leticia ranges from 66 to 78, with a mean of 

 73.75 and a standard deviation of 5.32; the number of such teeth in 

 three Trinidad specimens ranges from 51 to 56, with a mean of 54 and 

 a standard deviation of 2.65. 



Nomenclature. — Rivero (1961, p. 127) apparently was unaware that 

 the International Zoological Commission (1958) placed the trivial 

 name zonata Spix (as published in the combination Hyla zonata) on the 

 Official Index of Rejected Names in Zoology and suppressed for pur- 

 poses of priority both Acrodytes Fitzinger, 1843, and tibiatrix Lauren ti, 

 1768 (as published in the combination Hyla tibiatrix) ; and that it 



