330 U.S. NATIONAL MUSEUM BULLETIN 2 88 



ECUADOR: El Oro; 76 km. southeast of Buena vista, SUNHM 2569; Napo- 

 Pastaza, Montalvo, EPN 3007-8; Chicherota, EPN 2507-8; Avila, EPN 

 3013-4. 



Genus Phyllomedusa Wagler 



1830. Phyllomedusa Wagler, p. 201 (type species, Rana bicolor Boddaert). 



Generic diagnosis. — The frogs of this genus are characterized by 

 having the pupil vertical, the palpebral membrane not reticulate, 

 the tongue extensively free behind, and, in most species (all but 

 bicolor of the Colombian forms), the first toe longer than the second, 

 and in having very little or no webbing on the feet. 



Key to Colombian species of Phyllomedusa 



a 1 . First toe longer than second. 



b l . Posterior surface of thigh brightly patterned with black and white markings, 

 generally no dorsal pigment on tops of fingers. 

 c 1 . A distinct row of small white dots on each dorsolateral fold from behind 



eye to region of sacral hump P. blombergi 



c 2 . No row of distinct white spots along dorsolateral region from behind eye 



to region of sacral hump P. hypochondrialis hypochondrialis 



b 2 . Posterior surface of thigh plain or with a pattern but without conspicuous 



black and white markings, generally dorsal pigment on tops of fingers. 



c 1 . Upper surface of disks on fingers one and two without distinct white caps. 



d 1 . No series of rounded, distinct, white spots in the dorsal ground color 



along the sides between axilla and groin, or if such spots are present 



they are much smaller than tympanum. 



e l . A distinct pair of white spots below the vent P. nicefori 



e 2 . No distinct pair of white spots below the vent .... P. edentula 



d l . Some rather distinct, rounded, white spots in the dorsal ground color 



along the sides between axilla and groin, these spots about size of 



tympanum P. boliviana 



c 2 . Upper surface of disks of fingers one and two with distinct white caps. 



P. tarsius 

 a 2 . First toe shorter than second P. bicolor 



Phyllomedusa bicolor (Boddaert) 



Plate 40a-c 



1772. Rana bicolor Boddaert, p. 19 (type locality, Suriname) .—Shaw 1802, p. 126. 



1799. Calamita bicolor.— Schneider, 1799, p. 156.— Merrem, 1820, p. 170. 



1802. Hyla bicolor.— Baudin, 1802, p. 22; 1803, p. 40.— Spix, 1824, p. 42.— 



Fitzinger, 1826, p. 63. — Gravenhorst, 1829, p. 26.— Guerin-Meneville, 



1829, pi. 26, fig. 3.— Peters, 1873a, p. 220. 

 1830. Pyllomedusa bicolor.— Wagler, 1830, p. 201.— Tschudi, 1838, p. 70.— 



Dumeril and Bibron, 1841, p. 629.— Gunther, 1858, p. 120.— Boulenger, 



1882a, p. 427.— Beebe, 1919, p. 208.— Nieden, 1923, p. 337.— Miranda- 



Ribeiro, 1926, p. 103.— B. Lutz, 1950, p. 601.— Funkhouser, 1957, p. 



38.— Goin, 1958, p. 120.— Goin and Layne, 1958, p. 113.— Rivero, 1961, 



p. 147.— Gorham, 1963, p. 23. 

 1868. Phyllomedusa scleroderma Cope, p. 112 (type locality, Suriname). 



Diagnosis. — A large Phyllomedusa with the second toe longer than 



