430 U.S. NATIONAL MUSEUM BULLETIN 2 88 



color sometimes breaking up into two or three medium-sized spots on 

 the posterior femur, but usually in one large patch covering the entire 

 inner surface of the tibia, with a smaller spot on the upper tarsus. 

 The upper femoral surface has a wide longitudinal gray band sepa- 

 rating the two pink areas in front and behind, while the posterior 

 part of the belly is also pink. 



The adpressed heel reaches the anterior corner of the eye in about 

 45 percent of these frogs; between the eye and the nostril in 34 per- 

 cent; to the nostril in 8 percent; to the tip of the snout in 5 percent; 

 and to the center of the eye in 5 percent. The toes may lack webs or, 

 at most, have only slight traces of webs. A heel tubercle, present in 

 most specimens, ranges in size from a quite obvious, medium-sized, 

 pointed structure down to a series of several small ones; sometimes 

 it is a mere ridge across the heel. 



Specimens Examined 

 COLOMBIA 



Antioquia: Jeric6, MCZ 24895; Sons6n, AMNH 39327-8, 39330-7, 39340-2 

 Caldas: Laguneta, ANSP 25402; La Selva, Pueblo Rico, CNHM 54636, 

 54640-2, 54644, 54646-56, 54663, 54667, 54670, 54672-3, 54676-7, 54679, 

 54681, 54683, 54685, 54687-9, 54697-8, 54700, 54705, USNM 146969-70, 

 147233-9; Pueblo Rico, 5,200 ft., BM 1910.7.11.59-60; Santa Rosa de Aso, 

 AMNH 38818. 

 Cauca: Rio San Juan, USNM 147240-4; Quintana, near Popayan, CNHM 

 54414. 



Eleutherodactylus vertebralis (Boulenger) 



Plate 59d-i 



1886. Hylodes vertebralis Boulenger, 1886b, p. 415 (type locality, Intac, Ecua- 

 dor) .—Werner, 1901, p. 600.— Peracca, 1904, p. 26.— Nieden, 1923, p. 

 451. 



1934. Eleutherodactylus latidiscus (not of Boulenger). — Parker, 1934c, p. 267. 



1938. Eleutherodactylus vertebralis. — Parker, p. 440. — Gorham, 1963, p. 18. 



Description. — BM 78.1.25.15 (syntype), an adult female from Intac, 

 Ecuador. A pair of parallel, very slightly developed bony ridges on top 

 of head; frontoparietal area distinctly sunken. Vomerine teeth in two 

 small, widely separated, slanting series behind the choanae; tongue 

 one-half as wide as mouth opening, oval, its posterior border free and 

 unnotched; snout moderately long, slightly acuminate when viewed 

 from above, rounded in profile, the upper jaw extending well beyond 

 the lower. Nostrils dorsal, distinctly projecting, their distance from 

 end of snout one-half their distance from eye. Canthus rostralis dis- 

 tinct; loreal region very concave, flaring sharply outwards to the 

 upper lip. Eye large, prominent, its diameter three-fourths its distance 

 from tip of snout; interorbital diameter 1% times that of upper eyelid, 

 much greater than interval between nostrils. Tympanum distinct, its 



