FROGS OF COLOMBIA — COCHRAN AND GOIN 431 



greatest diameter one-half that of eye, separated from eye by an 

 interval equal to its own diameter. Fingers rather long, with prominent 

 lateral ridges, free, first finger much shorter than second, fourth longer 

 than either, reaching halfway on penultimate phalanx of third, the 

 disk of which covers the tympanum; a small oval thumb pad present; a 

 small double palmar callus; metacarpal tubercles well developed. Toes 

 moderately long, faintly webbed, third toe shorter than fifth, the disk 

 of which reaches to base of antepenultimate phalanx of fourth; disk of 

 fourth toe covering three-fourths the tympanum, and a little smaller 

 than that of third finger; a small oval inner metatarsal tubercle and a 

 smaller rounded outer one; a distinct tarsal ridge, ending near heel; a 

 heavy skinfold on heel and knee. Body rather stout, in postaxillary 

 region narrower than greatest width of head. When hind leg is ad- 

 pressed, heel reaches center of eye; when limbs are laid along the sides, 

 knee and elbow touch; when hind legs are bent at right angles of body, 

 heels overlap. Skin of upper parts pustular, with some very small 

 granules from behind eye along dorsolateral regions; a faint dorso- 

 lateral fold; sides finely to coarsely granular except in axilla and 

 groin; chin and chest smooth, belly coarsely granular; finer granules 

 around venter and on posterior femur; a heavy glandular ridge from 

 posterior corner of eye above tympanum, ending above shoulder; a 

 skinfold across the chest; traces of a ventral disk. 



Dimensions. — Head and body, 36 mm.; head length, 12.5 mm.; head 

 width, 13.5 mm.; femur, 15 mm.; tibia, 17 mm.; foot, 16.5 mm.; hand, 

 11.5 mm. 



Color in alcohol. — Dorsum uniform seal brown except for a wide 

 middorsal stripe from tip of snout to anus, ecru drab anteriorly to 

 vinaceous-cinnamon posteriorly; sides of body as well as anterior 

 femur cinnamon with a fine seal brown vermiculation becoming more 

 dense dorsally; posterior femur marbled with dark and light; sides of 

 head seal brown, the edge of the lip fawn color; venter ochraceous 

 with a few small scattered sepia spots on throat, chest and sides of 

 belly; palms of hands and soles of feet clay color, the disks and tuber- 

 cles drab, the outer digits darker. 



Remarks. — E. vertebralis may be grouped either with those species 

 having a sunken postoccipital area or with those having longitudinal 

 bony head ridges, since both features occur in this interesting frog. It 

 is actually more similar in appearance to latidiscus, which has the 

 depressed postoccipital region and no head ridges, than it is to cornutus, 

 which has heavy ridges; for this reason it is keyed near to latidiscus. 



A syntype of vertebralis was compared directly with a paratype of 

 the other large-disked frog, latidiscus. Both have a faint dorsolateral 

 fold. The vomerine teeth are small and slanting in vertebralis but are 

 heavy and transverse in latidiscus. The interorbital diameter is 1% 



