FROGS OF COLOMBIA — COCHRAN AND GOIN 433 



Eleutherodactylus f rater (Werner) 



Plate 54a-c 



1899. Hylodes frater Werner, p. 479 (type locality, Peperital, 1,000 m., near 



Villavicencio [Meta]).— Nieden, 1923, p. 442. 

 1963. Eleutherodactylus frater. — Gorham, 1963, p. 17. 



Description. — ZIMUG 67/235 (holotype), an adult from Peperital, 

 near Villavicencio, Meta, Colombia. No bony ridges on top of head; 

 frontoparietal area faintly depressed. Vomerine teeth in two very 

 weakly developed slanting series behind the choanae; tongue two- 

 thirds as wide as mouth opening, cordiform, its posterior border free 

 and deeply notched; snout broad, rounded when viewed from above 

 and in profile, the upper jaw extending slightly beyond the lower. 

 Nostrils dorsolateral, projecting, their distance from end of snout 

 one-half their distance from eye. Canthus rostralis curving, not very 

 distinct; loreal region faintly concave, sloping outwards to the upper 

 lip. Eye large, prominent, its diameter about four-fifths its distance 

 from tip of snout; interorbital diameter 1% times that of upper 

 eyelid, a little greater than interval between nostrils. Tympanum 

 small but distinct, its greatest diameter one-third that of eye, sepa- 

 rated from eye by an interval equal to its own diameter. Fingers 

 fairly long, with weak lateral ridges, free, first finger a little shorter 

 than second, fourth longer than either, reaching nearly to base of 

 disk of third, which covers three-fourths the tympanum; a prominent 

 oval thumb pad present; a weak, flat palmar callus; metacarpal 

 tubercles well developed. Toes moderately long, indistinctly webbed 

 at the base, third toe much shorter than fifth, the disk of which reaches 

 to base of antepenultimate phalanx of fourth; disk of fourth toe cover- 

 ing the tympanum, and a little larger than that of third finger; a 

 small oval inner metatarsal tubercle and an indistinct smaller outer 

 one; a weak inner tarsal ridge, ending near the knee; a heavy skinfold 

 on heel and knee. Body rather elongate, in postaxillary region nar- 

 rower than greatest width of head. When hind leg is adpressed, heel 

 reaches a point between eye and nostril; when limbs are laid along 

 the side, knee and elbow barely touch; when hind legs are bent at 

 right angles to body, heels overlap. Skin of upper parts pustular, 

 with larger granules and glands above the shoulder, on top of head, 

 and on sacrum; venter finely granular, the chin and chest smooth; 

 a thin, sharp glandular ridge from posterior corner of eye above 

 tympanum, ending behind corner of mouth; a heavy skinfold across 

 chest; a ventral disk; an external vocal sac in the male. 



Dimensions. — Head and body, 21.5 mm.; head length, 7.5 mm.; 

 head width, 8 mm.; femur, 8.5 mm.; tibia, 10.5 mm.; foot, 8 mm.; 

 hand, 5.5 mm. 



