FROGS OF COLOMBIA — COCHRAN AND GOIN 443 



1959. Eleutherodactylus cornutus. — Gin6s, 1959, p. 112, fig. 

 1961. Eleutherodactylus cornutus cornutus. — Rivero, 1961, p. 55. 



Description. — MCZ 19640, an adult from Pastaza River, Ecuador. 

 Heavy, smooth, bony ridges on top of head. Vomerine teeth in two 

 very heavy, transverse, arc-shaped, almost contiguous series behind 

 the choanae; tongue broadly cordiform, three-fifths as wide as mouth 

 opening, its posterior border free and sightly notched; snout broad 

 and rather short, rounded when viewed from above, slanting forward 

 in profile, the upper jaw extending a little beyond the lower, which 

 is obtusely pointed in front. Nostrils superolateral, projecting, their 

 distance from end of snout 1% times their distance from eye, separated 

 from each other by an interval less than their distance from eye. 

 Can thus rostralis rounded ; loreal region concave, the upper lip flaring 

 out below it. Eye rather small and deepset, its diameter about one-half 

 its distance from tip of snout; interorbital diameter almost twice that 

 of upper eyelid, slightly greater than interval between nostrils. Tym- 

 panum large, distinct, its greatest diameter five-sixths that of eye, 

 separated from eye by an interval equal to its own diameter. Fingers 

 moderate, with lateral ridges, free first finger much longer than sec- 

 ond, with bulbous tips instead of disks, a large oval thumb pad and a 

 flat palmar callus present; metacarpal tubercles distinct. Toes long, 

 free, third toe longer than fifth, with small but distinct disks, disk of 

 fourth toe covering about one-eighth the tympanum; a long oval inner 

 metatarsal tubercle and a very small round outer one; a heavy tarsal 

 ridge extending almost to heel; a distinct skinfold on heel and knee. 

 Body heavy and thickset, in postaxillary region narrower than the 

 unusually wide head. When hind leg is adpressed, heel reaches almost 

 to nostril; when limbs are laid along the sides, knee and elbow overlap; 

 when hind legs are bent at right angles to body, heels considerably 

 overlap. Dorsum heavily granular, with elongate glands on parietal 

 region near base of skull and on center of back; scattered large pustules 

 in rows across hind legs corresponding to the dark bands; venter 

 coarsely granular with somewhat smaller granules on chin; a pair of 

 heavy longitudinal ridges between the eyes; a small dermal horn on 

 upper eyelid; a narrow glandular ridge from posterior corner of eye 

 above tympanum, ending above shoulder; a faint skinfold across chest; 

 a ventral disk present. 



Dimensions. — Head and body, 56.5 mm.; head length, 21.5 mm.; 

 head width, 27 mm.; femur, 26.5 mm.; tibia, 29 mm.; foot, 27 mm.; 

 hand, 15 mm. 



Color in alcohol. — Dorsum drab, with a wide sepia interorbital 

 band preceded by a narrower pinkish buff band; clove brown spots 

 and short stripes emphasizing the glandular ridges on back; a sepia 



