446 U.S. NATIONAL MUSEUM BULLETIN 2 88 



becoming indistinct above venter; other warts prominent on dark 

 leg bands. Venter very finely granular, with heavy granulations on 

 femur around the vent; a weak glandular ridge from posterior corern 

 of eye above tympanum, a swollen indistinct parotoid gland above 

 and behind tympanum; a skinfold across the chest; no ventral disk; 

 in the male, a large median external vocal sac with many longitudinal 

 folds most prominent on the sides of the throat. 



Dimensions. — Head and body, 30 mm.; head length, 9.5 mm.; head 

 width, 10 mm.; femur, 12.5 mm.; tibia, 12 mm.; foot, 13 mm.; hand, 

 9.5 mm. 



Color in alcohol. — Dorsum sepia, with slightly darker areas around 

 the bases of the warts; side of body sepia to pale cinnamon below, with 

 several diagonal rows of small seal brown spots; side of head wood 

 brown with faint indications of a darker spot below eye; belly light 

 wood brown with several medium-sized clove brown spots; chest 

 russet; throat of male clove brown, the edges of the jaw russet; 

 limbs cinnamon with one or two darker crossbands; posterior femur 

 wood brown very slightly marbled with darker brown; lower surface 

 of femur pale wood brown with numerous small darker spots; anterior 

 femur pale cinnamon with slight continuations of the dark crossbands; 

 palms of hands and soles of feet cinnamon, their disks a little darker. 



Remarks. — The chief characters given in the original description of 

 ruthveni to separate it from pustulosus are the more granular belly, 

 which is immaculate posteriorly or marbled with small, light brown 

 spots, and the lack of large spots on the lower femoral surfaces. E. 

 pustulosus, on the contrary, was said to have a less granular belly with 

 larger dark coalescing spots, and large spots on the lower femur. The 

 difficulties are further increased by Dr. Netting's statement that 

 "Both E. pustulosus and E. ruthveni occur in Fundacion, Colombia 

 (the type locality of ruthveni), and the ranges of the two species may 

 be found to overlap throughout large areas in northern South 

 America." 



Graphs compiled from the critical measurements of Colombian frogs 

 with light-spotted or immaculate venters and others with dark- 

 spotted bellies show no significant differences in the body proportions 

 of the two groups. In a series of 27 specimens of pustulosus from El 

 Real, Darien Province, Panama, the belly is pale in 10 individuals, 

 dark in 13, and medium-shaded in four. Three of the dark-bellied 

 frogs had a light midventral line not invaded by the dark spotting; 

 two had only a few fairly large dark ventral spots, and eight had many 

 smaller ones. Considerable variation in the ventral coloration therefore 

 exists in a series of pustulosus from the same locality. 



As to the spotting of the lower femur, results are slightly more con- 

 clusive. In six Magdalena frogs considered to be "ruthveni" the lower 



