498 U.S. NATIONAL MUSEUM BULLETIN 2 88 



completely webbed, third and fifth subequal, disk of fourth toe equal 

 to diameter of eye; no distinct metatarsal tubercles; no tarsal ridge; 

 no dermal appendage on heel. Body not elongate, in postaxillary 

 region a little narrower than greatest width of head; when hind leg 

 is adpressed, heel reaches to eye; when limbs are laid along the side, 

 knee and elbow considerably overlap; when hind legs are bent at 

 right angles to body, heels just touch. Skin of upper parts highly 

 glandular, warty tubercles particularly conspicuous on snout, in- 

 cluding loreal and can thai regions, and on upper eyelids and occiput; 

 skin of throat, chest, belly, and lower femur uniformly granular. 

 A horizontal row of four distinct hyphen-shaped warts below the 

 vent. External vocal sacs present in the form of a thin, deflated sac 

 at the base of the angle of the jaw on each side. 



Dimensions. — Head and body, 74.2 mm.; head length, 22.5 mm.; 

 head width, 25.7 mm.; femur, 37.2 mm.; tibia, 38.5 mm.; heel-to-toe, 

 56.7 mm.; hand, 30 mm. 



Color in alcohol. — A uniform brown above and pale brown below. 

 Except as noted below, the dorsal and ventral surfaces of the limbs 

 are the same color as the dorsum and venter respectively. Except 

 for a few minute dark flecks in the sacral region of the back, the 

 pattern consists solely of dark inky markings of the left limbs. Four 

 narrow, transverse stripes cross the back of the hand just below the 

 wrist — these four stripes being so close together as to give the appear- 

 ance of a single dark band. On the left leg are a number of narrow, 

 transverse, inky stripes across the tops of the digits and the ankle 

 and a broader inky blob across the middle of the dorsal surface of 

 the shank. 



Variation. — Sexual dimorphism is noticeable. In the series of 

 specimens from Urrao (CNHM 69703-8, four males and two females), 

 the females not only are definitely shorter (60.4 and 61 mm.) than 

 the males (70.7, 74.2, 76.4, and 77 mm.) in head-and-body length 

 but are of a much lighter build. The diameter of the thigh in the fe- 

 males is no more than half the diameter of the thigh in the males, 

 and the diameter of the forearm is no more than one-fourth that of 

 the forearm in the males. A good idea of the size differential may be 

 gained by the fact that the two females average 15 grams each (just 

 as they are taken from the bottle) while the four males average 43 

 grams each. 



The most striking element of the pattern variation is its asym- 

 metry. In not a single specimen is the pattern symmetrical. The gen- 

 eral ground color in all except one male is reduced in places leaving 

 pale, washed-out patches of skin. In addition to this sort of ground 

 color variation three of the four males (but neither female) have dark 



