FROGS OF COLOMBIA — COCHRAN AND GOIN 417 



large wart between eyes; a faint dorsolateral fold; upper limb surfaces 

 fairly smooth; sides of body finely granular; venter nearly smooth 

 on chin and chest, finely granular and wrinkled transversely on 

 belly, granular on lower femur; a very heavy glandular ridge from 

 posterior corner of eye above tympanum, ending above shoulder; 

 a ridge along upper jaw, becoming more pronounced below eye; a short 

 heavy fold in front of shoulder; three skinfolds along sides of body 

 from above shoulder nearly to groin; a pronounced skinfold across 

 chest; a distinct ventral disk; a pair of external vocal sacs in the 

 male. 



Dimensions. — Head and body, 52 mm.; head length, 19 mm.; head 

 width, 20.5 mm.; femur, 21.5 mm.; tibia, 23 mm.; foot, 21 mm.; hand, 

 14 mm. 



Color in alcohol. — Dorsum seal brown, with the numerous glands 

 and tubercles raw umber; sides pale wood brown, with some narrow 

 dark and light diagonal fines on the posterior half to groin, and a few 

 scattered dark spots behind axilla; venter broccoli brown, the chin 

 with an irregular border of sepia marks separated by a short pale 

 line; throat and chest faintly dotted with sepia, belly heavily so; legs 

 with four wide, light-edged, clove brown crossbands, arms with two 

 less-apparent bands; posterior femur drab with a fine clove brown 

 vermiculation; sides of head clove brown with two light-edged sepia 

 spots below eye; palms of hands and soles of feet drab, their disks and 

 tubercles lighter. 



Remarks. — The leg is quite variable in length, as the adpressed heel 

 falls from the center of the tympanum to the tip of the snout, although 

 it reaches between the posterior and the anterior border of the eye in 

 66 percent of the frogs examined. The toes are distinctly webbed at 

 the base in about 70 percent and are very faintly webbed in the 

 remainder. A single strong tubercle is present on the heel of about 

 10 percent of the frogs, and in an equal number a flattened ridge or 

 skinfold occurs; in the great majority the heel has one or several 

 rather small tubercles or granules. 



Specimens CNHM 44119, 54312-3, and 54413 from Quintana, 

 Cauca, and USNM 152264 and 151270 from Purace, Cauca, resemble 

 the paratype BM 98.4.28.109 in essentials, as they have extremely 

 large disks on the third and fourth fingers, the same heavy canthus 

 with deeply concave loreal flaring almost horizontally toward the 

 upper lip, at least a trace of glandular dorsolateral ridges, and the 

 same diagonal lateral stripes and (usually) light-bordered crossbands 

 on the legs. But a great difference in development of the tubercles on 

 the head and body is apparent, as the interorbital tubercle, so obvious 

 in the paratype, may be represented by a small transverse ridge be- 

 tween the eyes, or that area may be nearly smooth. The tympanum 



