FROGS OF COLOMBIA — COCHRAN AND GOIN 453 



1799. Rana ocellata (not of Linnaeus). — Schneider, 1799, p. 117. — Daudin, 1S02, 

 p. 61, pi. 19; 1S03, p. 118.— Gravenhorst, 1829, p. 42. 



1830. Cystignalhus pachypus. — 'Wagler, 1830, p. 203 (part). 



1838. Cyslynathus ocellatus. — Tschudi, p. 78 (part). 



1S41. Cystignalhus ocellatus. — Dumeril and Bibron, p. 396 (part). — De Betta, 

 1876, p. 695. 



1865. Gnalhophysa labryrinthica. — Cope, p. 112. 



1865. Gnathophysa ocellata.— Cope, 1865b, p. 112; 1866a, p. 73. 



1873. Cystignathus pentadactylus. — Peters, 1873a, p. 198. 



1875. Leptodaclylus goliath Jiminez de la Espada, p. 57 (type localities, Quijos, 

 Archidona, Chuntambo, Sierra de Guacamayos, eastern Ecuador). 



1875. Leptodactylus stenodema Jiminez de la Espada, p. 64 (type locality, San 

 Jose de Moti, Canton de Quijos, Ecuador). 



1882. Leptoiaclylus pentadactylus. — Boulenger, 1882a, p. 241 (part); 1898a, p. 

 122; 1913, p. 1025— Giinther, 1901, p. 212 (part) .— Peracca, 1904, 

 p. 34.— Barbour, 1914, p. 251.— Baumann, 1917, pp. 132, 138.— Beebe, 

 1919, p. 209; 1925, p. 123.— Ruthven, 1922, p. 54.— Nieden, 1923, p. 

 473 (part).— Breder, 1925, p. 328; 1927, p. 72; 1946, p. 402.— L. Miiller, 

 1927, p. 259.— A. Lutz, 1927, p. 40.— Ditmars, 1929, p. 53.— Crawford, 

 1931, p. 30.— Crawford and Jones, 1933, p. 88.— Hellmich, 1939, p. 393, 

 fig. 12.— Dunn, 1944, p. 511.— H. Travassos, 1947, pp. 619-627.— B. 

 Lutz, 1948, p. 30.— Valente, 1949, p. 335.— Schreiber and Velucci, 1951, 

 p. 73.— Houssay, 1952, p. 137.— Johnson, 1952, p. 284.— Oltmanns, 

 1952, p. 370.— Bhaduri, 1953, p. 35, pi. 9, figs. 52, 53.— Vinton, 1951, 

 p. 73.— Cei, 1953, p. 510; 1956, p. 43, figs. 58, 59.— Brattstrom and 

 Howell, 1954, p. 116.— GinSs, 1959, p. 114.— Goin, 1959, p. 136.— 

 Fouquette, 1960d, pp. 201, 209, fig. 11, table 2.— Cochran, 1961, p. 62, 

 pi. 26.— Gallardo, 1961b, p. 34.— Gorham, 1933, p. 19. 



1927. Leptodactylus pentadactylus pentadactylus. — L. Miiller, p. 279. — Parker, 

 1935, p. 507.— Andersson, 1945, p. 47, fig. 16.— Lutz and Kloss, 1952, 

 p. 640.— Goin and Layne, 1958, p. 100.— Rivero, 1961, p. 35; 1964a, 

 p. 298. 



Description. — USNM 147053, an adult female from Leticia, Ama- 

 zonas, Colombia. Vomerine teeth in two heavy, A -shaped, narrowly 

 separated series between the choanae; tongue large, two-thirds as wide 

 as mouth opening, cordiform, its posterior border free and deeply 

 notched; snout moderate, rounded when viewed from above, sloping 

 towards the lip border in profile, the upper jaw extending well beyond 

 the lower. Nostrils dorsolateral, scarcely projecting, their distance from 

 end of snout two-third their distance from eye. Canthus rostralis 

 strongly defined; loreal region concave, sloping to the upper lip. Eye 

 large, prominent, its diameter two-thirds its distance from tip of 

 snout; interorbital diameter equal to that of upper eyelid, equal to 

 interval between nostrils. Tympanum distinct, its greatest diameter 

 three-fourths that of eye, separated from eye by an interval equal to 

 two-thirds its own diameter. Fingers moderate, with heavy lateral 

 ridges, free, first finger longer than second and fourth and reaching 

 halfway on ultimate phalanx of third; an oval thumb pad present; a 



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