FISHES OF THE PHILIPPINE ISLANDS AND ADJACENT SEAS 667 



ANALYSIS OF GENERA 



a\ Engraxjunae. Body moderately long ; anal free from strongly forked 

 caudal ; upper pectoral rays not extended as slender filaments. 

 b^. Silvery coloration absent only on back. 

 c\ Body more or less compressed; abdominal scutes at least between pec- 

 torals and anal. 

 d\ Upper pectoral ray not produced ; A. 30 to 50, length 3 to 5 in total 

 length without caudal. 

 e\ Teeth in jaws partly caninelike; scales 42 to 50. 



f. Dorsal origin behind anal origin Lycothrissa 



f. Dorsal origin well before anal origin Xenengraulis 



e^ Teeth uniform, minute ; scales 35 to 45 ; dorsal origin before anal 



origin Thrissocles 



rfl Upper pectoral ray produced ; A. 50 to 75, fin 2 to 2% in total length 



without caudal Setipinna 



c". Body more or less robust, partly cylindrical ; dentition feeble, without 

 canines ; no abdominal scutes ; fins without produced rays ; A. 17 to 



23 Engraulis 



h^. Silvery coloration limited chiefly to a conspicuous lateral band ; abdominal 



scutes largely or only between pectorals and ventrals Anchoviella 



a". CoiLiiNAE. Body greatly elongate, tail long and tapering; caudal little or 

 not foiked, lower portion continuous with long anal ; upper pectoral rays 

 produced as slender filaments Coilia 



Genus LYCOTHRISSA Gunther 



Lycothrissa GtJNTHEE, Cat. Fishes British Mus., vol. 7, p. 399, 1868. (Type, 

 Engraulis crocodilus Bleeker, monotypic.) 



Body elongate, compressed. Head small. Snout short. Eye 

 small. Mouth oblique. Maxillary narrow, extending to mandibular 

 joint, supplementary bone only partly ossified. Jaws with caninelike 

 teeth, especially few in mandible. Small teeth on vomer, palatines, 

 pterygoids, and tongue. Scales moderate, thin, deciduous, with irreg- 

 ular curved transverse lines. Not very conspicuous abdominal scutes 

 from below pectorals to vent. Dorsal origin behind anal origin. 

 Anal very long. Upper pectoral ray not produced. Ventral small. 



One species. 



LYCOTHRISSA CROCODILUS (Bleeker) 



Engraulis crocodilus Blexkeb, Nat. Tijdschr. Nederland. Indie, vol. 1, p. 15, 

 1851 (type locality: Banjermassing, in rivers); Verb. Batav. Genootsch. 

 (Harding.), vol. 24, p. 35, 1852 (Borneo). — GiJNTHER, Cat. Fishes British 

 Mus., vol. 7, p. 399, 1868 (type).— Sauvage, Bull. Soc. Philom. Paris, ser. 

 7, vol. 7, p. 151, 1883 (Maenam). — Tibant, Service Oceauogr. Peches Indo- 

 Chine, 6" note, pp. 114, 174, 1929 (Phuoc Hai, Thudaumot). 



Engraulis {Lycothrissa) crocodilus Bleeker, Atlas Ichth. Ind. Neerland., vol. 6, 

 pi. (10)208, fig. 1, 1866-72.— Mabtens, Preuss. Exped. Ost-Asien, vol. 1, p. 

 404, 1876 (Bangkok; Pontianak and Sinkawang, Borneo). 



Lycothrissa crocodilus Bleeker, Atlas Ichth. Ind. Neerland., vol. 6, p. 125, 1866-72 

 (Borneo). — Weber and Be,v.ufort, Fishes Indo-Australian Archipelago, vol. 

 2, p. 31, fig. 16, 1913 (Djambi). — Chabanaud, Service Oceanogr. Peches Indo- 

 Chine, 1* note, p. 8, 1926 (Cambodia). — Fowler, Proc. Acad, Nat. Sci. Phila- 



